Revision of the Dysmorphoptilidae (Hemiptera: Cicadomorpha: Prosboloidea) of the Queensland Triassic — Part 2
Author
Lambkin, Kevin J.
text
Zootaxa
2016
4092
2
207
218
journal article
10.11646/zootaxa.4092.2.4
0847a84a-637d-4368-8bd8-0eb7171e5f59
1175-5326
267859
56072014-E67D-4866-A131-6693C2836B5E
Eoscarterella
Evans, 1956
Type
species.
Eoscarterella media
Evans, 1956, by original designation.
Diagnostic characters.
Tegmen approximately
10 mm
long, asymmetrically lobate, apical margin at RA2 gently convex, at M strongly convex, anterior margin fairly straight; precostal carina and marginal membrane very narrow; hypocostal carina well-developed, extending along most of costal space (Figs 11,12); sculpture comprising irregular, short, longitudinally orientated rugae, particularly coarse in basal costal space, grading to a finely punctate sculpture apically (
Fig.12
); strigil not detected in basal costal space; hyposubcostal carina short, angulately concave; clavus not preserved; primary fork of R proximal to clavus ½ length; primary fork of RA proximal to apex of clavus, at about level of CuA fork; stem of RA1 directed anteriorly, more or less in line with stem of RA, entering costal margin at about ⅔ tegmen length, simple or with 1 or 2 anterior branches; RA2 quite extensive, occupying a large area antero-apically, with 3-5 more or less pectinate branches, the middle one forked; RA2 and
RP
joined by a crossvein;
RP
simple, not up-curved apically; M1+2 simple, M3+4 deeply forked; M and CuA joined basally by a weak crossvein; apical m-cu upright; CuA1 strongly concave; CuA2 straight, much shorter than CuA1.
Notes.
The retention of a well-developed hypocostal carina is unique among Australian dysmorphoptilids and is considered plesiomorphic for the family (Dmitry Shcherbakov, pers com.). The peculiar tegmen shape and surface sculpture, and the branching pattern of M (M1+2 simple, M3+ 4 forked) are considered as the apomorphies for the genus. It is similar to
Eoscartoides
, the Australian Permian
Belmontocarta
Evans (Evans 1958) and the unnamed Triassic species from
Kyrgyzstan
illustrated by Shcherbakov and Popov (2002, fig.184) in sharing the evenly rounded tegmen without an emargination, the early entry of RA1 into the costal margin rather than the apex and the associated extensive and antero-apically positioned RA2.