Diversity and taxonomic review of Leptusa Kraatz (Coleoptera: Staphylinidae: Aleocharinae) from Great Smoky Mountains National Park, U. S. A., with descriptions of four new species Author Park, Jong-Seok Author Carlton, Christopher E. Author Ferro, Michael L. text Zootaxa 2010 2662 1 27 journal article 10.5281/zenodo.199052 a07f4b3c-e602-4b4e-95dc-87fef597d912 1175-5326 199052 Subgenus Adoxopisalia Pace, 1989 Leptusa ( Adoxopisalia Pace, 1989 : 130 ); Klimaszewski et al. , 2004 : 23 ; Gouix & Klimaszewski, 2007 : 61 . Type species. Leptusa opaca Casey, 1893 ; designated by Pace (1989) . Diagnosis. See Pace (1989) and Klimaszewski et al. (2004) .