Revision of some types of Philonthus Stephens and Gabrius Stephens from southern South America (Coleoptera: Staphylinidae), with nomenclatural notes
Author
Maus, Mariana Chani-Posse De
text
Zootaxa
2009
2034
31
42
journal article
10.5281/zenodo.186351
bb17e039-5346-4e01-a9ba-59b87e9e6a4c
1175-5326
186351
Philonthus cribriventris
Bernhauer, 1912
(
Figs 10, 11
)
Philonthus cribriventris
Bernhauer, 1912
: 43
;
Bernhauer & Schubert, 1914
: 334
(catalog);
Bruch, 1915
: 444
(catalog);
Blackwelder, 1944
: 132
(list);
Herman, 2001
: 2793
(catalog); Chani-Posse, 2004: 230 (list).
Philonthus catamarcanus
var.
densior
Bernhauer, 1916
: 36
;
Scheerpeltz, 1933
: 1335
(cat.),
syn. nov.
Type
material.
Lectotype
of
Philonthus cribriventris
(here designated;
Figs. 10
a–b), 3, with labels: “Rep.
Argentina
/ Prov. Tucuman /
30.I.1908
/ C. Bruch”, “
cribriventris
Brnh
/
Typus
”, “Chicago NHMus. / M. Bernhauer Collection”, “
Lectotype
Philonthus cribriventris
Bernhauer, 1912
/ Des. Chani-Posse de Maus 2008” (
FMNH
).
Paralectotype
:
3, with labels: “Rep.
Argentina
/ Prov. Tucuman /
30.I.1908
”, “
cribriventris
Brnh
/ Cotypus”, “Chicago NHMus. / M. Bernhauer Collection”, “
Paralectotype
Philonthus cribriventris
Bernhauer, 1912
/ Des. Chani-Posse de Maus 2008” (
FMNH
).
Holotype
of
Philonthus catamarcanus
var.
densior
(
Fig. 11
): Ƥ, “Rep.
Argentina
/ Prov. Tucuman / 19 / Richter”, “
v.
densior
Bernh.
/
Typus
unicus”, “Chicago NHMus. / Bernhauer Collection” (
FMNH
).
Additional material.
ARGENTINA
: Tucumán:
3, “Rep.
Argentina
/ Prov. Tucuman / 19 / Richter”, “in Bernhauer handwriting "
longicornis
Gr.
" / det. Bernhauer”, “
v.
densior
Bernh.
/ det. Bernhauer”, “Chicago NHMus. / Bernhauer Collection” (
FMNH
).
Diagnosis.
Philonthus cribriventris
may be identified by the eyes shorter than temples, the first segment of metatarsus longer than last segment, the basal area of abdominal terga with coarse punctuation, and the apex of tergum 10 subacute in both sexes.
Notes.
In the original description of
P. cribriventris
,
Benhauer (1912)
stated that two specimens were available for the description and gave two different
type
localities with different collector names: “
Argentinien
, Prov. Tucumán,
19. / I. 1908
(C. Bruch, H. Richter), Chaunar Region (Jensen)”. Both specimens listed above agree with the description by
Bernhauer (1912)
, and both have one same locality data and collection date. One of them includes the name of one of the collectors for the given locality. I have no way to decide which specimen may not belong to the original series. Therefore I have decided to consider both of them part of the
type
series, and designate the specimen with the collector´s name on its label as
lectotype
.
Bernhauer (1916)
described
P. catamarcanus
var.
densior
as a variety (“Abart”) from Tucuman province, while the
type
locality of
P. catamarcanus
was Catamarca and San Luis provinces. According to Article 45.6 (
ICZN 1999
) the name
densior
is of subspecific rank and available. Bernhauer explicitely stated that the
type
series of
P. catamarcanus
var.
densior
included a single specimen, but did not mention whether this was a male or a female. Considering that the female specimen of
P. catamarcanus
var.
densior
listed above is labeled as “
Typus
unicus”, and locality label corresponds to the locality mentioned in the description, this female specimen is regarded as the
holotype
. The male specimen from the same locality listed in Additional material was not labeled by Bernhauer as the
type
, and therefore cannot be the
holotype
of
P. catamarcanus
var.
densior
.
Comments.
Bernhauer (1916: 36)
described
P. densior
as a variety of
catamarcanus
and since then it was a valid taxon at the subspecific rank. The
type
material of
P. densior
(
lectotype
Ƥ,
paralectotype
3) (
Fig. 11
) agrees with the
lectotype
of
P. cribriventris
(
Figs. 10
a–b) in external characters and in the shape of the aedeagus. Therefore,
P. d e n s i o r
is removed from synonymy with
P. catamarcanus
(at the rank of species) and placed in synonymy with
P. cribriventris
Bernhauer, 1912
.