<document id="74C73C12362ACC36B95FAFF575833EC4" ID-CLB-Dataset="4031" ID-DOI="10.5852/ejt.2021.750.1365" ID-GBIF-Dataset="818d0bc4-31a6-4319-86a3-6e6b16ab72df" ID-ISSN="2118-9773" ID-Zenodo-Dep="5451747" ID-ZooBank="CE5D8517-AC04-4AF9-BD78-A2D6E21D33A8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630516667691" checkinUser="felipe" docAuthor="Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z. &amp; Ahrens, Dirk" docDate="2021" docId="038B976CFF9FFFFACAEBCA4F179CCB53" docLanguage="en" docName="ejt-750_94-123.pdf" docOrigin="European Journal of Taxonomy 750 (750)" docSource="http://dx.doi.org/10.5852/ejt.2021.750.1365" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Sayloria Frey 1973" docType="treatment" docVersion="9" lastPageNumber="114" masterDocId="FFB2EF14FF8BFFEEC84EC925155FCE04" masterDocTitle="Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor" masterLastPageNumber="123" masterPageNumber="94" pageNumber="114" updateTime="1699290068827" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="E12175F9F82BE7A7577C6B45030A7C07">Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor</mods:title>
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<mods:namePart id="E1A9DC5F622907519161431080EFAC98">Pacheco, Thaynara L.</mods:namePart>
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<mods:affiliation id="4D2E8CF101C5345B7CD057CB41825026">Laboratório de Coleoptera, Departamento de Entomologia, Museu Nacional, Universidade Federal do Rio de Janeiro, Rio de Janeiro, Brazil. &amp; Centre of Taxonomy and Evolutionary Research, Zoologisches Forschungsmuseum Alexander Koenig, Bonn, Germany.</mods:affiliation>
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<mods:namePart id="FBA11B63AEB355DFE75058A2C9DBB677">Monné, Marcela L.</mods:namePart>
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<mods:affiliation id="A19E2C9ADE6C9013D9B29846D186A533">Laboratório de Coleoptera, Departamento de Entomologia, Museu Nacional, Universidade Federal do Rio de Janeiro, Rio de Janeiro, Brazil. &amp; Laboratório de Scarabaeoidologia, Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, Cuiabá, Brazil.</mods:affiliation>
<mods:nameIdentifier id="041EA1E57DB2AA545D3BC909BE316B4C" type="email">mlmonne2@gmail.com&amp;vazdemello@gmail.com</mods:nameIdentifier>
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<mods:namePart id="2743081C27C7C2BFA0C8887B85570F4B">Vaz-De, Fernando Z.</mods:namePart>
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<mods:affiliation id="C40A578DF8288AB147C784A911C6518E">Laboratório de Scarabaeoidologia, Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, Cuiabá, Brazil.</mods:affiliation>
<mods:nameIdentifier id="564C3DE388583B00101072913276A1AF" type="email">vazdemello@gmail.com</mods:nameIdentifier>
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<mods:namePart id="8A0979874CF564D5B789B021EA432A27">Ahrens, Dirk</mods:namePart>
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<mods:affiliation id="B0D046BFBB7A7515812D82D752CF3904">Centre of Taxonomy and Evolutionary Research, Zoologisches Forschungsmuseum Alexander Koenig, Bonn, Germany.</mods:affiliation>
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<taxonomicName id="4C225DF9FF9FFFFACAEBCA4F16D0CD81" authority="Frey, 1973" authorityName="Frey" authorityYear="1973" box="[677,911,874,901]" class="Insecta" family="Melolonthidae" genus="Sayloria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="114" phylum="Arthropoda" rank="genus">
<emphasis id="B956FA68FF9FFFFACAEBCA4F1655CD81" bold="true" box="[677,778,874,901]" italics="true" pageId="20" pageNumber="114">Sayloria</emphasis>
<bibRefCitation id="EFB35B8BFF9FFFFACB5FCA4E16D0CD81" author="Frey G." box="[785,911,875,901]" pageId="20" pageNumber="114" pagination="315 - 366" refId="ref13316" refString="Frey G. 1973. Synopsis der sudamerikanischen Sericinen. Entomologische Arbeiten aus dem Museum Frey 1973: 315 - 366." type="journal article" year="1973">Frey, 1973</bibRefCitation>
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<taxonomicName id="4C225DF9FF9FFFFAC8F3CA8814FACDC3" authority="Frey, 1973" authorityName="Frey" authorityYear="1973" box="[189,421,941,967]" class="Insecta" family="Melolonthidae" genus="Sayloria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="114" phylum="Arthropoda" rank="genus">
<emphasis id="B956FA68FF9FFFFAC8F3CA88147FCDC3" box="[189,288,941,967]" italics="true" pageId="20" pageNumber="114">Sayloria</emphasis>
<bibRefCitation id="EFB35B8BFF9FFFFAC966CA8814FACDC3" author="Frey G." box="[296,421,941,967]" pageId="20" pageNumber="114" pagination="315 - 366" refId="ref13316" refString="Frey G. 1973. Synopsis der sudamerikanischen Sericinen. Entomologische Arbeiten aus dem Museum Frey 1973: 315 - 366." type="journal article" year="1973">Frey, 1973</bibRefCitation>
</taxonomicName>
(nec 
<bibRefCitation id="EFB35B8BFF9FFFFAC9A8CA8817E5CDC3" author="Chalumeau F." box="[486,698,941,967]" pageId="20" pageNumber="114" pagination="137 - 142" refId="ref12816" refString="Chalumeau F. 1981. Nouveaux genre et sous-genres d'Eupariini (Aphodiinae) d'Amerique tropicale. Nouvelle Revue d'Entomologie 11: 137 - 142." type="journal article" year="1981">Chalumeau, 1981</bibRefCitation>
): 324 (as subgenus of 
<taxonomicName id="4C225DF9FF9FFFFACBF1CA881176CDC3" authorityName="Erichson" authorityYear="1835" box="[959,1065,941,967]" class="Insecta" family="Melolonthidae" genus="Symmela" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="114" phylum="Arthropoda" rank="genus">
<emphasis id="B956FA68FF9FFFFACBF1CA881176CDC3" box="[959,1065,941,967]" italics="true" pageId="20" pageNumber="114">Symmela</emphasis>
</taxonomicName>
).
</paragraph>
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<treatmentCitationGroup id="AB320154FF9FFFFAC8F3CACA16D8CA2A" pageId="20" pageNumber="114">
<taxonomicName id="4C225DF9FF9FFFFAC8F3CACA147FCA0D" authorityName="Frey" authorityYear="1973" box="[189,288,1007,1033]" class="Insecta" family="Melolonthidae" genus="Sayloria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="114" phylum="Arthropoda" rank="genus">
<emphasis id="B956FA68FF9FFFFAC8F3CACA147FCA0D" box="[189,288,1007,1033]" italics="true" pageId="20" pageNumber="114">Sayloria</emphasis>
</taxonomicName>
(as subgenus of 
<taxonomicName id="4C225DF9FF9FFFFAC9A5CACA1783CA0D" authority="Frey, 1973" authorityName="Frey" authorityYear="1973" box="[491,732,1007,1034]" class="Insecta" family="Melolonthidae" genus="Symmela" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="114" phylum="Arthropoda" rank="genus">
<emphasis id="B956FA68FF9FFFFAC9A5CACA170ACA0D" box="[491,597,1007,1033]" italics="true" pageId="20" pageNumber="114">Symmela</emphasis>
<bibRefCitation id="EFB35B8BFF9FFFFACA12CAD51783CA0D" author="Frey G." box="[604,732,1007,1034]" pageId="20" pageNumber="114" pagination="315 - 366" refId="ref13316" refString="Frey G. 1973. Synopsis der sudamerikanischen Sericinen. Entomologische Arbeiten aus dem Museum Frey 1973: 315 - 366." type="journal article" year="1973">Frey, 1973</bibRefCitation>
</taxonomicName>
) – 
<treatmentCitation id="0A83006BFF9FFFFACB4ACAD516E3CA0D" author="Evans A. V." box="[772,956,1007,1034]" page="47" pageId="20" pageNumber="114" year="2003">
<bibRefCitation id="EFB35B8BFF9FFFFACB4ACAD516E3CA0D" author="Evans A. V." box="[772,956,1007,1034]" pageId="20" pageNumber="114" pagination="1 - 458" refId="ref13003" refString="Evans A. V. 2003. A checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa 211: 1 - 458. https: // doi. org / 10.11646 / zootaxa. 211.1.1" type="journal article" year="2003">Evans 2003: 47</bibRefCitation>
</treatmentCitation>
(checklist). — Evans &amp; 
<treatmentCitation id="0A83006BFF9FFFFACCABCACA1458CA29" author="Smith A. B. T. &amp; Evans A. V." page="29" pageId="20" pageNumber="114" year="2005">
<bibRefCitation id="EFB35B8BFF9FFFFACCABCACA1458CA29" author="Smith A. B. T. &amp; Evans A. V." pageId="20" pageNumber="114" pagination="29 - 60" refId="ref14061" refString="Smith A. B. T. &amp; Evans A. V. 2005. A supplement to the checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae) with notes on their tribal classification. Zootaxa 1032: 29 - 60. https: // doi. org / 10.11646 / zootaxa. 1032.1.2" type="journal article" year="2005">Smith 2005: 29</bibRefCitation>
</treatmentCitation>
(checklist); 2007: 29 (checklist); 2009: 30 (checklist).
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</paragraph>
</subSubSection>
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<heading id="D0D59116FF9FFFFAC8F3CD701407CA6B" bold="true" box="[189,344,1109,1135]" fontSize="11" level="3" pageId="20" pageNumber="114" reason="3">
<emphasis id="B956FA68FF9FFFFAC8F3CD701407CA6B" bold="true" box="[189,344,1109,1135]" pageId="20" pageNumber="114">
<typeStatus id="549998D8FF9FFFFAC8F3CD7015A5CA6B" box="[189,250,1109,1135]" pageId="20" pageNumber="114">Type</typeStatus>
species
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9D267AFF9FFFFAC8F3CDA0165ECA9B" blockId="20.[189,769,1156,1183]" box="[189,769,1156,1183]" pageId="20" pageNumber="114">
<taxonomicName id="4C225DF9FF9FFFFAC8F3CDA0171CCA9A" authority="Saylor, 1946" authorityName="L. W. Saylor" authorityYear="1946" box="[189,579,1156,1183]" class="Insecta" family="Melolonthidae" genus="Astaena" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="114" phylum="Arthropoda" rank="species" species="bicoloripes">
<emphasis id="B956FA68FF9FFFFAC8F3CDA014F8CA9A" box="[189,423,1156,1183]" italics="true" pageId="20" pageNumber="114">Astaena bicoloripes</emphasis>
<bibRefCitation id="EFB35B8BFF9FFFFAC9E0CDA1171CCA9A" author="Saylor L. W." box="[430,579,1156,1182]" pageId="20" pageNumber="114" pagination="215 - 232" refId="ref13932" refString="Saylor L. W. 1946. New South American scarab beetles of the genus Astaena. Revista de Entomologia 17: 215 - 232." type="journal article" year="1946">Saylor, 1946</bibRefCitation>
</taxonomicName>
(by monotypy).
</paragraph>
</subSubSection>
<subSubSection id="C33875F1FF9FFFFAC8F3CDE3179CCB53" pageId="20" pageNumber="114" type="diagnosis">
<paragraph id="8B9D267AFF9FFFFAC8F3CDE31469CAE4" blockId="20.[189,310,1222,1248]" box="[189,310,1222,1248]" pageId="20" pageNumber="114">
<heading id="D0D59116FF9FFFFAC8F3CDE31469CAE4" bold="true" box="[189,310,1222,1248]" fontSize="11" level="3" pageId="20" pageNumber="114" reason="3">
<emphasis id="B956FA68FF9FFFFAC8F3CDE31469CAE4" bold="true" box="[189,310,1222,1248]" pageId="20" pageNumber="114">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B9D267AFF9FFFFAC8F3CDD3179CCB53" blockId="20.[189,1398,1269,1368]" pageId="20" pageNumber="114">Brown, reddish-brown, blackish coloration, often bicolored. Antennae with eight antennomeres; antennal club of males with three antennomeres. Abdomen without a robust lateral carina. Mentum deeply sinuated medially at anterior margin.</paragraph>
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</treatment>
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