Reassessing the phylogeny and divergence times of sloths (Mammalia: Pilosa: Folivora), exploring alternative morphological partitioning and dating models Author Casali, Daniel M Author Boscaini, Alberto Author Gaudin, Timothy J Author Perini, Fernando A text Zoological Journal of the Linnean Society 2022 2022-12-01 196 4 1505 1551 https://academic.oup.com/zoolinnean/article/196/4/1505/6617197 journal article 10.1093/zoolinnean/zlac041 0024-4082 7381236 Urumacotheriinae PP = 60, age = 17.82 Mya (14.03–21.51). This clade was originally proposed with fewer taxa ( Negri & Ferigolo, 2004 ), but was expanded here to include both Urumacotherium and Pseudoprepotherium . It did not receive high node support, despite being present in trees from all analyses. Urumacotheriinae was supported by seven synapomorphies (five for both methods and two exclusively for BI): molariform morphology of Cf1/ cf1 (BI); posterior edge of mandibular condyle inclined posterodorsally; eversion of lateral edge of symphyseal spout (BI); premolariform portion of palate roughly equal or longer than the length of molariform toothrow; presence of a distinct neck at the base of occipital condyles; occipital condyles elongated anteroposteriorly in ventral view; and lesser tubercle of humerus proximally projected as much as greater tubercle.