Revision of Coelana Kramer, 1964 (Hemiptera, Cicadellidae, Neocoelidiinae) and taxonomic notes on the genus
Author
Marques-Costa, Ana Paula
Author
Cavichioli, Rodney Ramiro
text
Zootaxa
2007
1547
33
42
journal article
10.5281/zenodo.178025
0c048217-17f9-4784-b2dd-cb8b92e1f76a
1175-5326
178025
Coelana
Kramer, 1964
(
Figures 1–29
)
Coelidiana
(
Coelana
)
DeLong, 1953
: 95
(key to genera) [
nomen nudum
];
Metcalf, 1964
: 112
(catalogue, citation as an invalid name).
Type
species not designated.
Coelana
DeLong
;
Oman
et al.
, 1990
: 201
, 305 (catalogue, citation as an invalid name);
Freytag & Sharkey, 2002
: 254
(citation, number of species);
Dietrich, 2003
: 701
(citation, distribution).
Coelana
Kramer, 1964
: 261
, 269–270 (key to genera, description, key to species).
Type
species:
Neocoelidia modesta
Baker
, 1898
. Designated by
Kramer, 1964
: 269
.
Diagnosis.
Color stramineous to yellow with small black spot at apex of crown (
Figs. 1
,
10
); carina present in transition between crown and frons; posterior margin of pronotum emarginated, broadly indented (
Figs. 1
,
10
); forewing with venation indistinct, except for claval suture and apical cells (
Figs. 4
,
13
); male genitalia: pygofer greatly inflated, with large ventral hook, a pair of long inner processes dorsally, and laterally with a long ventrodorsal suture (
Figs. 5
,
14
); connective Y-shaped (
Figs. 8
,
17
) and aedeagus simple, without processes (
Figs. 9
,
18
); anal tube with a pair of ventral processes (
Figs. 5
,
14
).
Description.
In dorsal view, body approximately cylindrical, not flat.
Head. Crown almost quadrangular, slightly projected anteriorly and slightly wider than long; median length of crown approximately one half of transocular width and approximately equal to interocular width; anterior margin broadly angular, with carina in the transition between crown and frons; lateral margins, adjacent to compound eyes, elevated (not at the same level of the dorsum of compound eyes) and carinate; surface flat, smooth, without grooves, with weak coronal maculae at the base of crown, close to compound eyes; coronal suture indistinct (
Figs. 1
,
10
); ocelli located on anterior margin of head, in the transition between crown and frons, closer to compound eyes than to midline, above antennal ledges; in lateral view, antennal ledges oblique and carinate (
Figs. 3
,
12
); antennae setaceous, scape and pedicel enlarged, flagellum longer than basal half of forewings (
Figs. 2–3
,
11–12
); frons with length approximately two times basal width, profile convex, muscle impressions generally indistinct, not swollen on the central portion of base; frontogenal sutures reaching ocelli; apical margin of maxillas not extending beyond apex of clypeus; lorum small, approximately halfmoon shaped, inferior margin not reaching apical margin of clypeus; gena partially concealing proepisternum, only apex visible in anterior view; epistomal suture distinct and complete, slightly curved; clypeus almost rectangular, lateral margins parallel, generally without a gibbosity in lateral view, and apical margin straight (
Figs. 2–3
,
11–12
).
Thorax. Pronotum slightly wider than head, width between humeri slightly larger than transocular width; median length approximately one third of width between humeri; lateral margins rounded and posterior margin emarginated, broadly indented (
Figs. 1
,
10
); dorsal surface with transverse grooves; dorsopleural carina complete and evident; in ventral view, mesothorax moderately swollen; scutellum (
sensu
Young 1968
) as long as maximum width, with distinct preapical fold (
Figs. 1
,
10
). Forewing (
Figs. 4
,
13
) with rounded apex, without punctuations, hyaline, about 3–3.5 times longer than greatest width; venation indistinct, except for claval suture and apical cells; four apical cells, all almost rectangular, first larger than second, third slightly enlarged apically; second and fourth apical cells approximately same distance to base, closer to forewing base than third cell; anteapical cells distinct or not; appendix narrow, extending only along first apical cell. Hindwing with three apical cells, R4+5 and M1+2 apically convergent, fused, forming a single vein. Hindleg: femoral setal formula 2+2+0; tibial row PD with long setae little separated amongst themselves, distributed throughout length of tibia, intercalary setae absent; row AD with long and robust setae in form of spines, with two or more small intercalary setae present; row AV with setae moderately long, distributed only on apical two thirds, intercalary setae absent; row PV densely bristly, with short setae at base, becoming progressively longer toward apex; first tarsomere as long as combined length of two distal ones, plantar surface with two parallel rows of short setae.
Abdomen. Male genitalia: pygofer greatly inflated, longer than subgenital plates, with large ventral hook, a pair of long inner processes dorsally, and laterally with a long suture, extending from ventral margin to near dorsal margin (
Figs. 5
,
14
); subgenital valve entirely fused to subgenital plates; subgenital plates fused on the basal and median thirds, separated only on apical third (
Figs. 6
,
15
); styles simple, not bifurcated, with sclerotized hook-like apex, curved ventrally (
Figs. 7–8
,
16–17
); connective Y-shaped, articulated to base of aedeagus (
Figs. 8
,
17
); aedeagus simple, without processes (
Figs. 9
,
18
); anal tube with a pair of ventral processes (
Figs. 5
,
14
).
Total length (body + forewings). 6.0–
7.5 mm
.
Geographical distribution.
Argentina
,
Bolivia
and
Brazil
.
Comments.
This genus is similar to
Coelidiana
Oman
regarding the external morphology and general color of the species. It possesses some diagnostic characters that easily separate it from the other genera: general color stramineous to yellow with a small black spot at apex of crown and the characteristic male genitalia, mainly the morphology of the pygofer and anal tube.