Documenting diversity in the Amazonian butterfly genus Bia (Lepidoptera, Nymphalidae) Author Penz, Carla M. Author Casagrande, Mirna M. Author Devries, Phil Author Simonsen, Thomas J. text Zootaxa 2017 4258 3 201 237 journal article 33146 10.11646/zootaxa.4258.3.1 97942778-cef4-4cf6-b4e4-c077e633daa8 1175-5326 569729 DE89E162-B255-474C-A824-20DFA551DBBD Bia actorion (Linnaeus) ( Figs 1a , 3 , 11 a–b, 12) Type species. Diagnostic re-description. General characters as described for the genus. (1) MF DFW white apical ocelli small to large. (2) MF DFW orange band varies from thin ( Venezuela , Alto Orinoco) to medium-wide; orange scales extended proximally along veins, including the anterior tip of discal cell in some specimens ( Brazil , Amazonas , Manaus, but not all specimens). (3) M DFW iridescent band from anal margin to approximately half of the CuA2 cell (one specimen from Venezuela with DFW iridescence clearly reaching CuA2); F DFW iridescent band well developed but variable; usually broad below CuA2, and more extended towards tornus than those of female decaerulea decaerulea and rebeli . (4) M DFW androconial organ on the CuA-CuA1-CuA2 intersection dark brown, matching scale color of surrounding area. This character is shared by B. actorion and B. decaerulea . (5) M DHW discal androconial pad varies from being nearly black and darker than associated hairpencil ( Venezuela ; Guyana ; Suriname ; Brazil , Manaus) to dark brown and of similar color to hairpencil (photos from Colombian specimens, Ecuador , northern Peru ; Brazil , northern Pará ) . (6) M DHW discal hairpencil dark brown. (7) F VFW ripple pattern of the postmedial area slightly less dense than that of M in some localities ( Brazil and Guyana ) and clearly less dense in others ( Venezuela , Ecuador ). Male genitalia ( Figs 2 and 11 a): tip of valva usually narrow, spiny projection usually of medium height, infrequently tall, spines larger in specimens from the Guyana region ; gnathos wide (Guyana region) to narrow (other localities). Type material. The iconotype designated by Hemming (1964) from Clerck’s Icones ( Fig. 1a ) does not show the color of androconial organs and also lacks locality data, both of which are important attributes for definitive identification of actorion . We therefore recognize that a neotype should be selected for this species, which will be done separately from this study. Note that specimens in the Linnean Collection are not types (male http://linneanonline.org/14787/ and female http://linnean-online.org/14788/ last accessed 30 August 2016 ; see also http:// butterfliesofamerica.com/L/t/ Bia _a.htm last accessed 30 August 2016 ). Remarks. Geographical variation in some diagnostic characters serves as the basis for the subspecific classification proposed below.