<document id="21EB79DCC8BC678820DFBAF4D76039C3" ID-CLB-Dataset="157784" ID-DOI="10.18590/euscorpius.2019.vol2019.iss281.1" ID-GBIF-Dataset="67120029-de04-4f6e-b3e8-460ef3896e70" ID-ISSN="1536-9307" ID-Zenodo-Dep="7143610" ID-ZooBank="FEBA0106-02A3-4465-8D39-9FF32634EEF" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638261886193" checkinUser="felipe" docAuthor="Lowe, Graeme &amp; Kovařík, František" docDate="2019" docId="03BB8789FF8FFF992463D371FB2EFF87" docLanguage="en" docName="Euscorpius.281.1-128.pdf" docOrigin="Euscorpius 281" docSource="https://doi.org/10.18590/euscorpius.2019.vol2019.iss281.1" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Teruelius mahafaliensis Lowe &amp; Kovařík 2019, comb. n." docType="treatment" docVersion="7" lastPageNumber="61" masterDocId="FF82FFF1FFB1FFA72751D179FFB2FC46" masterDocTitle="Review of Grosphus Simon 1880 with description of Teruelius gen n a new buthid genus from Madagascar (Scorpiones Buthidae)" masterLastPageNumber="128" masterPageNumber="1" pageNumber="61" updateTime="1699471319803" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="68612E93D356D0C835D93D6525554BBC">Review of Grosphus Simon 1880 with description of Teruelius gen n a new buthid genus from Madagascar (Scorpiones Buthidae)</mods:title>
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<mods:namePart id="98B3C6B8154707CFCB876A58591EDA7E">Lowe, Graeme</mods:namePart>
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<treatment id="03BB8789FF8FFF992463D371FB2EFF87" ID-DOI="http://doi.org/10.5281/zenodo.7143722" ID-GBIF-Taxon="202133055" ID-Zenodo-Dep="7143722" LSID="urn:lsid:plazi:treatment:03BB8789FF8FFF992463D371FB2EFF87" httpUri="http://treatment.plazi.org/id/03BB8789FF8FFF992463D371FB2EFF87" lastPageNumber="61" pageId="62" pageNumber="61">
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<taxonomicName id="4C124D1CFF8FFF992463D371FA87FE67" ID-CoL="B87TV" authority="(Lourenco et al., 2004)" authorityName="Lowe &amp; Kovařík" authorityYear="2019" baseAuthorityName="Lourenco et al." baseAuthorityYear="2004" box="[818,1333,520,545]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="62" pageNumber="61" phylum="Arthropoda" rank="species" species="mahafaliensis" status="comb. nov.">
<emphasis id="B966EA8DFF8FFF992463D371FB88FE66" bold="true" box="[818,1082,520,544]" italics="true" pageId="62" pageNumber="61">Teruelius mahafaliensis</emphasis>
(Lourenço et al., 2004)
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<emphasis id="B966EA8DFF8FFF99226DD370FAC8FE67" bold="true" box="[1340,1402,521,545]" pageId="62" pageNumber="61">comb</emphasis>
. 
<emphasis id="B966EA8DFF8FFF9922D9D370FA25FE67" bold="true" box="[1416,1431,521,545]" pageId="62" pageNumber="61">n</emphasis>
.
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<paragraph id="8BAD369FFF8FFF992463D350FB2EFF87" blockId="62.[818,1476,520,961]" pageId="62" pageNumber="61">
<materialsCitation id="3B7A3CC2FF8FFF992463D350FC28FEE7" ID-GBIF-Occurrence="3920707317" collectingDate="2004-09-08" collectionCode="ZMUH" collectorName="W. R. Lourenco" location="NE Manandona" pageId="62" pageNumber="61" specimenCount="1" specimenCount-male="1" stateProvince="Central Region" typeStatus="holotype">
<specimenCount id="9D14FD16FF8FFF992463D350FCE6FE07" box="[818,852,553,577]" pageId="62" pageNumber="61" type="male">1♂</specimenCount>
(
<typeStatus id="54A9883DFF8FFF992434D350FC76FE07" box="[869,964,553,577]" pageId="62" pageNumber="61" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C124D1CFF8FFF9924BFD350FB2BFE06" authorityName="Lourenco" authorityYear="2013" box="[1006,1177,553,577]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="62" pageNumber="61" phylum="Arthropoda" rank="species" species="rossii">
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<originalDetermination id="A80C8E63FF8FFF9924BFD350FB2BFE06" box="[1006,1177,553,577]" pageId="62" pageNumber="61">Grosphus rossii</originalDetermination>
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<figureCitation id="13292A1AFF8FFF9923FBD350FAFBFE07" box="[1194,1353,553,577]" captionStart="Figures 522–525" captionStartId="106.[113,190,1981,2003]" captionTargetBox="[150,1435,241,1931]" captionTargetId="figure-39@106.[127,1460,232,1934]" captionTargetPageId="106" captionText="Figures 522–525. Teruelius mahafaliensis. Habitus. Male holotype of G. rossii (522–523) and male from near the type locality of T. mahafaliensis (524–525), dorsal (522, 524) and ventral (523, 525) views. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7143817" httpUri="https://zenodo.org/record/7143817/files/figure.png" pageId="62" pageNumber="61">Figs. 522–523</figureCitation>
, 
<figureCitation id="13292A1AFF8FFF992206D350FA0AFE07" box="[1367,1464,553,577]" captionStart-0="Figures 526–537" captionStart-1="Figures 538–541" captionStartId-0="107.[113,190,1928,1950]" captionStartId-1="108.[113,190,1981,2003]" captionTargetBox-0="[113,1458,234,1912]" captionTargetBox-1="[129,1451,241,1964]" captionTargetId-0="figure-80@107.[113,1468,234,1919]" captionTargetId-1="figure-37@108.[123,1459,234,1977]" captionTargetPageId-0="107" captionTargetPageId-1="108" captionText-0="Figures 526–537. Teruelius mahafaliensis, female (526, 529–530, 535), male (527–528, 531–534) and male holotype of G. rossii (536–537). Sternopectinal region (526–527). Carapace and tergites I–II (528). Pedipalp chela, movable finger dentate margin (529). Pedipalp chela dorsal (530–531) and external (532) views. Pedipalp patella dorsal (533) view. Pedipalp femur and trochanter dorsal (534) view. Metasoma V and telson lateral (535) view. Metasoma and telson dorsal (536) and ventral (537) views. Scale bar: 10 mm (536–537)." captionText-1="Figures 538–541. Teruelius mahafaliensis, male holotype of G. rossii. Left legs tibia, basitarsus and telotarsus, retrolateral views. Leg I (538), leg II (539), leg. III (540), leg IV (541)." figureDoi-0="http://doi.org/10.5281/zenodo.7143819" figureDoi-1="http://doi.org/10.5281/zenodo.7143821" httpUri-0="https://zenodo.org/record/7143819/files/figure.png" httpUri-1="https://zenodo.org/record/7143821/files/figure.png" pageId="62" pageNumber="61">536–541</figureCitation>
), 
<collectingRegion id="49D6F87DFF8FFF992463D330FC68FE27" box="[818,986,585,609]" country="Ghana" name="Central" pageId="62" pageNumber="61">Central Region</collectingRegion>
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<location id="8ECD6044FF8FFF9924B6D330FB24FE27" LSID="urn:lsid:plazi:treatment:03BB8789FF8FFF992463D371FB2EFF87:8ECD6044FF8FFF9924B6D330FB24FE27" box="[999,1174,585,609]" name="NE Manandona" pageId="62" pageNumber="61" stateProvince="Central Region">NE Manandona</location>
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<location id="8ECD6044FF8FFF9923F2D330FAF1FE27" LSID="urn:lsid:plazi:treatment:03BB8789FF8FFF992463D371FB2EFF87:8ECD6044FF8FFF9923F2D330FAF1FE27" box="[1187,1347,585,609]" name="S of Antsirabe" pageId="62" pageNumber="61" stateProvince="Central Region">S of Antsirabe</location>
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<date id="FFAC105FFF8FFF992200D330FA0DFE27" box="[1361,1471,585,609]" pageId="62" pageNumber="61" value="2004-09-08">
<collectingDate id="EFE8E9B7FF8FFF992200D330FA0DFE27" box="[1361,1471,585,609]" pageId="62" pageNumber="61" value="2004-09-08">8.IX.2004</collectingDate>
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, leg. 
<collectorName id="26E75349FF8FFF992435D310FBA7FEC6" box="[868,1045,617,640]" pageId="62" pageNumber="61">W. R. Lourenço</collectorName>
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<materialsCitation id="3B7A3CC2FF8FFF9924F2D3F0FB17FF67" ID-GBIF-Occurrence="3920707355" collectingDate="2005-02-20" collectionCode="FMNH" collectorName="Mahafaly Plateau" elevation="120" latitude="-24.736668" location="10.5 km SE Itampolo" longLatPrecision="92" longitude="44.023167" pageId="62" pageNumber="61" specimenCode="FMNH 73598" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Toliara">
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<figureCitation id="13292A1AFF8FFF9924A7D3F0FBE3FEE7" box="[1014,1105,649,673]" captionStart="Figures 9–20" captionStartId="11.[116,190,1913,1935]" captionTargetBox="[129,1461,231,1903]" captionTargetId="figure-144@11.[129,1461,231,1903]" captionTargetPageId="11" captionText="Figures 9–20. Position of femur trichobothrium d 2in Grosphus and Teruelius gen. n. Dorsal surfaces of proximal pedipalp femur of adult females (♀, left panels) and adult males (♂, right panels), underUV fluorescence to highlightgranulation,carinae and trichobothrial areolae.Whitearrows indicate positions of trichobothrium d in each image. G. madagascariensis (9, 10; 2 samples from Anjiro (9) and Andasibe (10) show consistency of d 2 2 position),G. hirtus(11), G. voahangyae (12), T. ankarafantsika (13, 15; 2 samples fromAmpijoroa show consistency ofd 2position),T. ankarana (14), T. flavopiceus (16), T. grandidieri (17), T. limbatus (18), T. mahafaliensis (19), and T. olgae (20). Scale bars: 1 mm (9–20♀) or 500 μm (20 ♂)." figureDoi="http://doi.org/10.5281/zenodo.7143618" httpUri="https://zenodo.org/record/7143618/files/figure.png" pageId="62" pageNumber="61">Figs. 19</figureCitation>
, 
<figureCitation id="13292A1AFF8FFF99230FD3F0FBC9FEE7" box="[1118,1147,649,673]" captionStart="Figures 40–51" captionStartId="22.[113,188,1930,1952]" captionTargetBox="[113,1474,234,1920]" captionTargetId="figure-124@22.[113,1474,234,1922]" captionTargetPageId="22" captionText="Figures 40–51. Female basal pectinal teeth in Grosphus and Teruelius gen. n. Ventral views of proximal left pectine of females shown under UV fluorescence to highlight cuticular surface texture, setation and absence of peg sensillae on basal tooth vs. their presence on other teeth. G. sp. nr hirtus (40), G. madagascariensis (41), G. hirtus (42), G. voahangyae (43), T. ankarafantsika (44–45; 2 samples fromAmpijoroa show variation in tooth shape), T. ankarana (46), T. flavopiceus (47), T. grandidieri (48), T. limbatus (49), T. mahafaliensis (50), and T. olgae (51). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7143634" httpUri="https://zenodo.org/record/7143634/files/figure.png" pageId="62" pageNumber="61">50</figureCitation>
, 
<figureCitation id="13292A1AFF8FFF9923D9D3F0FB17FEE7" box="[1160,1189,649,673]" captionStart="Figures 71–85" captionStartId="26.[113,190,1701,1723]" captionTargetBox="[149,1434,231,1678]" captionTargetId="figure-287@26.[146,1446,225,1690]" captionTargetPageId="26" captionText="Figures 71–85. Hemispermatophores and capsule regions of Teruelius gen n. Multi-panel figures show: whole hemispermatophore; whole hemispermatophore and/or capsule with flagellum; capsule region in convex (or convex compressed), anterior and posterior views (panels in left to right sequence). Right hemispermatophores. Figure 71. T. ankarana, whole hemispermatophore. Scale bar: 4 mm. Figure 72. T. ankarana, capsule, left mirrored, Forêt d’Ankavanana, FMNH. Scale bar: 500 μm. Figure 73. T. grandidieri, whole hemispermatophore (flagellum truncated). Scale bar: 4 mm. Figure 74. T. grandidieri, capsule, Antsakabe River, FMNH. Scale bar: 500 μm. Figure 75. T. flavopiceus, whole hemispermatophore. Scale bar: 2 mm. Figure 76. T. flavopiceus, capsule, Madagascar, GLPC, FKCP. Scale bar: 500 μm. Figure 77. T. annulatus, capsule, Tsimanampetsotsa National Park, GLPC, FKCP. Scale bar: 500 μm. Figure 78. T. ankarafantsika, capsule and flagellum. Scale bar: 400 μm. Figure 79. T. ankarafantsika, capsule, Forêt d’Ankavanana, FMNH. Scale bar: 200 μm. Figure 80. T. ankarafantsika, capsule, Réserve Forestière de l’Ankarafantsika, FMNH. Scale bar: 200 μm. Figure 81. T. olgae, capsule, Itampolo village, FMNH. Scale bar: 500 μm. Figure 82. T. limbatus, whole hemispermatophore (flagellum truncated). Scale bar: 2 mm. Figure 83. T. limbatus, capsule, Forêt d’Ianasana, FMNH. Scale bar: 500 μm. Figure 84. T. mahafaliensis, capsule views, Zombitse-Vohibasia National Park, GLPC, FKCP. Scale bar: 500 μm. Figure 85. T. intertidalis, capsule, Madagascar, GLPC, FKCP. Scale bar: 500 μm." figureDoi="http://doi.org/10.5281/zenodo.7143641" httpUri="https://zenodo.org/record/7143641/files/figure.png" pageId="62" pageNumber="61">84</figureCitation>
, 
<figureCitation id="13292A1AFF8FFF9923E3D3F0FB6EFEE7" box="[1202,1244,649,673]" captionStart="Figures 95–110" captionStartId="18.[113,190,1408,1430]" captionTargetBox="[113,1475,255,1894]" captionTargetId="figure-131@18.[113,1474,255,1915]" captionTargetPageId="18" captionText="Figures 95–110: Orthochirus mesopotamicus comb. n., from Iraq, Basra (ca. 30°48’N 47°34’E). Figures 95, 97, 100–110. Male, carapace and tergites I–III (95), sternopectinal region and sternites (97). Pedipalp chela, dorsal (100), external (101), and ventrointernal (102) views. Pedipalp patella, dorsal (103), external (104), and ventral (105) views. Pedipalp femur and trochanter, internodorsal (106) and dorsoexternal (107) views. Pedipalp chela, movable finger dentate margin (108). Distal segments of right legs III–IV, retrolateral views (109–110). Figures 96, 98–99. Female, carapace and tergites I–III (96), sternopectinal region and sternites (98), and pedipalp chela dorsoexternal (99). The trichobothrial pattern is indicated in Figures 101–104, 106–107 (white circles)." pageId="62" pageNumber="61">104</figureCitation>
, 
<figureCitation id="13292A1AFF8FFF9923B8D3F0FAA1FEE7" box="[1257,1299,649,673]" captionStart="Figures 127–132" captionStartId="39.[113,190,1954,1976]" captionTargetBox="[132,1462,256,1937]" captionTargetId="figure-68@39.[113,1474,231,1947]" captionTargetPageId="39" captionText="Figures 127–132. Ventral aspect of metasoma and telson of adult males (♂) or females (♀) of representative Teruelius gen. n under UV fluorescence to reveal carination and granulation. T. ankarafantsika (127), T. ankarana (128), T. flavopiceus (129), T. grandidieri (130), T. mahafaliensis (131), and T. olgae (132). Scale bars: 4 mm." figureDoi="http://doi.org/10.5281/zenodo.7143675" httpUri="https://zenodo.org/record/7143675/files/figure.png" pageId="62" pageNumber="61">131</figureCitation>
, 
<figureCitation id="13292A1AFF8FFF992271D3F0FAF8FEE7" box="[1312,1354,649,673]" captionStart="Figures 133–144" captionStartId="40.[113,190,1293,1315]" captionTargetBox="[118,1474,249,1273]" captionTargetId="figure-523@40.[113,1474,227,1273]" captionTargetPageId="40" captionText="Figures 133–144. Ventral setation of telotarsus III in Grosphus and Teruelius gen. n. Ventral surfaces of right telotarsus III of adult males (♂) or females (♀), shown under UV fluorescence to highlight setation. Macrosetae appear dark with strongly fluorescent cuticular sockets at their base. Putative chemosensory microsetae appear bright. G. hirtus (= G. h. garciai) (133), G. hirtus (134), G. madagascariensis (135, from Anjiro, 136 from Andasibe), G. voahangyae (137), T. ankarafantsika (138), T. limbatus (139), T. mahafaliensis (140), T. ankarana (141), T. grandidieri (142), T. flavopiceus (143), and T. olgae (144). Scale bars: 500 μm (133–140, 144) and 1 mm (141–143)." figureDoi="http://doi.org/10.5281/zenodo.7143677" httpUri="https://zenodo.org/record/7143677/files/figure.png" pageId="62" pageNumber="61">140</figureCitation>
, 
<figureCitation id="13292A1AFF8FFF992206D3F0FA33FEE7" box="[1367,1409,649,673]" captionStart="Figures 145–157" captionStartId="41.[113,190,1502,1524]" captionTargetBox="[113,1474,234,1490]" captionTargetId="figure-392@41.[113,1474,234,1490]" captionTargetPageId="41" captionText="Figures 145–157. Comparative intensity of UV fluorescence in Grosphus and Teruelius gen. n. Photographic comparison of fluorescence emission intensities of representative species of each genus, including adult males (♂) or females (♀). G. madagascariensis (145 from Anjiro, 148 from Andasibe), G. hirtus (146), G. voahangyae (147), G. hirtus (= G. h. garciai) (149), T. ankarafantsika (150), T. grandidieri (151), T. olgae (152), T. flavopiceus (153), T. ankarana (154–155), T. mahafaliensis (156), and T. limbatus (157). Images acquired under identical intensities of UV excitation (395 nm LED source, 0.35 A current) and camera exposure (Canon EOS 7D Mark II, 100 mm f/13, 0.5 s, ISO 320), with 475 nm longpass filter to block excitation wavelengths. Scale bars: 10 mm (145–150, 152–153, 156–157), 20 mm (151, 154–155)." figureDoi="http://doi.org/10.5281/zenodo.7143681" httpUri="https://zenodo.org/record/7143681/files/figure.png" pageId="62" pageNumber="61">156</figureCitation>
, 
<figureCitation id="13292A1AFF8FFF9922DED3F0FCEEFE87" captionStart="Figures 165–180" captionStartId="63.[113,190,1958,1980]" captionTargetBox="[113,1473,312,1921]" captionTargetId="figure-73@63.[113,1473,312,1921]" captionTargetPageId="63" captionText="Figures 165–180. Carapace of representative Grosphus and Teruelius gen. n. G. madagascariensis (165, 168), G. hirtus (166–167), G. voahangyae (169), T. ankarafantsika (170–171), T. ankarana (172), T. flavopiceus (173–174), T. grandidieri (175), T. limbatus (176), T.mahafaliensis (177–178) andT. olgae (179–180).UV fluorescence,♂male, ♀female.Scale bars:2mm (165–171, 176–180),4 mm (172–175)." figureDoi="http://doi.org/10.5281/zenodo.7143694" httpUri="https://zenodo.org/record/7143694/files/figure.png" pageId="62" pageNumber="61">177– 178</figureCitation>
, 
<figureCitation id="13292A1AFF8FFF99243BD3D0FC27FE87" box="[874,917,681,705]" captionStart="Figures 181–195" captionStartId="64.[113,190,1953,1975]" captionTargetId="figure-71@64.[113,1474,242,1670]" captionTargetPageId="64" captionText="Figures 181–195. Telson of representativeGrosphus andTeruelius gen. n. G. hirtus (181–182), G. madagascariensis (183–184), G. voahangyae (185), T. ankarafantsika (186–187), T. ankarana (188), T. flavopiceus (189), T. grandidieri (190–191), T. limbatus (192), T. mahafaliensis (193) and T. olgae (194–195). UV fluorescence, ♂ male, ♀ female. Scale bars: 2 mm (181–187, 192–195), 4 mm (188–191)." figureDoi="http://doi.org/10.5281/zenodo.7143700" httpUri="https://zenodo.org/record/7143700/files/figure.png" pageId="62" pageNumber="61">193</figureCitation>
), 
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, 
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<quantity id="4CEA9B7AFF8FFF99233FD3D0FB7AFE87" box="[1134,1224,681,705]" metricMagnitude="4" metricUnit="m" metricValue="1.05" pageId="62" pageNumber="61" unit="km" value="10.5">10.5 km</quantity>
SE Itampolo
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(village), 
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a. s. l.
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, 
<date id="FFAC105FFF8FFF9923EFD3B0FA82FEA7" box="[1214,1328,713,737]" pageId="62" pageNumber="61" value="2005-02-20">
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, 
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in disturbed spiny bush on 
<collectorName id="26E75349FF8FFF992365D390FB43FF47" box="[1076,1265,745,769]" pageId="62" pageNumber="61">Mahafaly Plateau</collectorName>
, leg. 
<collectionCode id="ED03AE5AFF8FFF992279D390FA89FF46" box="[1320,1339,745,768]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="62" pageNumber="61" type="Museum">V</collectionCode>
. 
<location id="8ECD6044FF8FFF992214D390FA76FF47" LSID="urn:lsid:plazi:treatment:03BB8789FF8FFF992463D371FB2EFF87:8ECD6044FF8FFF992214D390FA76FF47" box="[1349,1476,745,769]" name="Soarimalala" pageId="62" pageNumber="61" stateProvince="Toliara">Soarimalala</location>
&amp; 
<collectorName id="26E75349FF8FFF99241CD270FC47FF67" box="[845,1013,777,801]" pageId="62" pageNumber="61">S. M. Goodman</collectorName>
(
<specimenCode id="DBB49EE4FF8FFF992353D270FB2FFF67" box="[1026,1181,777,801]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="62" pageNumber="61" type="Museum">FMNH 73598</specimenCode>
)
</materialsCitation>
; 
<materialsCitation id="3B7A3CC2FF8FFF9923FED270FC30FFC7" ID-GBIF-Occurrence="3920707307" collectionCode="FKCP, GLPC" elevation="692" latitude="-22.683533" location="Isoky" longLatPrecision="1" longitude="44.86392" municipality="Zombitse-Vohibasia" pageId="62" pageNumber="61" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Toliara" typeStatus="holotype">
<specimenCount id="9D14FD16FF8FFF9923FED270FB64FF67" box="[1199,1238,777,801]" pageId="62" pageNumber="61" type="male">2♂</specimenCount>
<specimenCount id="9D14FD16FF8FFF992386D270FB43FF67" box="[1239,1265,777,801]" pageId="62" pageNumber="61" type="female">1♀</specimenCount>
(
<figureCitation id="13292A1AFF8FFF9923AED270FAE5FF67" box="[1279,1367,777,801]" captionStart="Figures 71–85" captionStartId="26.[113,190,1701,1723]" captionTargetBox="[149,1434,231,1678]" captionTargetId="figure-287@26.[146,1446,225,1690]" captionTargetPageId="26" captionText="Figures 71–85. Hemispermatophores and capsule regions of Teruelius gen n. Multi-panel figures show: whole hemispermatophore; whole hemispermatophore and/or capsule with flagellum; capsule region in convex (or convex compressed), anterior and posterior views (panels in left to right sequence). Right hemispermatophores. Figure 71. T. ankarana, whole hemispermatophore. Scale bar: 4 mm. Figure 72. T. ankarana, capsule, left mirrored, Forêt d’Ankavanana, FMNH. Scale bar: 500 μm. Figure 73. T. grandidieri, whole hemispermatophore (flagellum truncated). Scale bar: 4 mm. Figure 74. T. grandidieri, capsule, Antsakabe River, FMNH. Scale bar: 500 μm. Figure 75. T. flavopiceus, whole hemispermatophore. Scale bar: 2 mm. Figure 76. T. flavopiceus, capsule, Madagascar, GLPC, FKCP. Scale bar: 500 μm. Figure 77. T. annulatus, capsule, Tsimanampetsotsa National Park, GLPC, FKCP. Scale bar: 500 μm. Figure 78. T. ankarafantsika, capsule and flagellum. Scale bar: 400 μm. Figure 79. T. ankarafantsika, capsule, Forêt d’Ankavanana, FMNH. Scale bar: 200 μm. Figure 80. T. ankarafantsika, capsule, Réserve Forestière de l’Ankarafantsika, FMNH. Scale bar: 200 μm. Figure 81. T. olgae, capsule, Itampolo village, FMNH. Scale bar: 500 μm. Figure 82. T. limbatus, whole hemispermatophore (flagellum truncated). Scale bar: 2 mm. Figure 83. T. limbatus, capsule, Forêt d’Ianasana, FMNH. Scale bar: 500 μm. Figure 84. T. mahafaliensis, capsule views, Zombitse-Vohibasia National Park, GLPC, FKCP. Scale bar: 500 μm. Figure 85. T. intertidalis, capsule, Madagascar, GLPC, FKCP. Scale bar: 500 μm." figureDoi="http://doi.org/10.5281/zenodo.7143641" httpUri="https://zenodo.org/record/7143641/files/figure.png" pageId="62" pageNumber="61">Figs. 84</figureCitation>
, 
<figureCitation id="13292A1AFF8FFF992230D270FA3EFF67" box="[1377,1420,777,801]" captionStart="Figures 196–210" captionStartId="65.[113,190,1950,1972]" captionTargetId="figure-77@65.[113,1474,234,1589]" captionTargetPageId="65" captionText="Figures 196–210. Female basal pectinal teeth of representative Grosphus and Teruelius gen. n. G. hirtus (196–198), G. madagascariensis (199), G. madagascariensis, paratype of G. mandena(200), T. ankarafantsika (201),T. ankarana (202), T. annulatus (203),T. bistriatus (204), T. flavopiceus (205), T. grandidieri (206), T. intertidalis (207), T. limbatus (208), T. mahafaliensis (209), T. feti (210)." figureDoi="http://doi.org/10.5281/zenodo.7143706" httpUri="https://zenodo.org/record/7143706/files/figure.png" pageId="62" pageNumber="61">209</figureCitation>
, 
<figureCitation id="13292A1AFF8FFF9922C4D270FA72FF67" box="[1429,1472,777,801]" captionStart="Figures 211–226" captionStartId="66.[113,190,1920,1942]" captionTargetBox="[117,1466,234,1743]" captionTargetId="figure-88@66.[113,1474,234,1743]" captionTargetPageId="66" captionText="Figures 211–226. Ventral tarsal setation of legs III or IV in Grosphus and Teruelius gen. n. G. goudoti (211), G. hirtus (212), G. hirtus (G. garciai) (213), G. madagascariensis (214), G. madagascariensis (G. mandena) (215), Teruelius ankarana (216), T. flavopiceus (217), T. annulatus (218), T. ankarafantsika (219–220), T. bistriatus (221), T. intertidalis (222), T. grandidieri (223), T. mahafaliensis (224), T. limbatus (225), T. feti (226)." figureDoi="http://doi.org/10.5281/zenodo.7143708" httpUri="https://zenodo.org/record/7143708/files/figure.png" pageId="62" pageNumber="61">224</figureCitation>
, 
<figureCitation id="13292A1AFF8FFF992463D250FC26FF07" box="[818,916,809,833]" captionStart-0="Figures 522–525" captionStart-1="Figures 526–537" captionStartId-0="106.[113,190,1981,2003]" captionStartId-1="107.[113,190,1928,1950]" captionTargetBox-0="[150,1435,241,1931]" captionTargetBox-1="[113,1458,234,1912]" captionTargetId-0="figure-39@106.[127,1460,232,1934]" captionTargetId-1="figure-80@107.[113,1468,234,1919]" captionTargetPageId-0="106" captionTargetPageId-1="107" captionText-0="Figures 522–525. Teruelius mahafaliensis. Habitus. Male holotype of G. rossii (522–523) and male from near the type locality of T. mahafaliensis (524–525), dorsal (522, 524) and ventral (523, 525) views. Scale bar: 10 mm." captionText-1="Figures 526–537. Teruelius mahafaliensis, female (526, 529–530, 535), male (527–528, 531–534) and male holotype of G. rossii (536–537). Sternopectinal region (526–527). Carapace and tergites I–II (528). Pedipalp chela, movable finger dentate margin (529). Pedipalp chela dorsal (530–531) and external (532) views. Pedipalp patella dorsal (533) view. Pedipalp femur and trochanter dorsal (534) view. Metasoma V and telson lateral (535) view. Metasoma and telson dorsal (536) and ventral (537) views. Scale bar: 10 mm (536–537)." figureDoi-0="http://doi.org/10.5281/zenodo.7143817" figureDoi-1="http://doi.org/10.5281/zenodo.7143819" httpUri-0="https://zenodo.org/record/7143817/files/figure.png" httpUri-1="https://zenodo.org/record/7143819/files/figure.png" pageId="62" pageNumber="61">524–535</figureCitation>
), 
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, 
<collectingMunicipality id="6BC9ACE5FF8FFF99230CD250FA84FF07" box="[1117,1334,809,833]" pageId="62" pageNumber="61">Zombitse-Vohibasia</collectingMunicipality>
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forrest margin, 
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a. s. l.
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, 
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, (
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<specimenCount id="9D14FD16FF8FFF9924C8D210FC70FFC7" box="[921,962,873,897]" pageId="62" pageNumber="61" type="male">4♂</specimenCount>
<specimenCount id="9D14FD16FF8FFF992493D210FC6FFFC7" box="[962,989,873,897]" pageId="62" pageNumber="61" type="female">1♀</specimenCount>
, 
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, 
<collectingCounty id="62CC4E13FF8FFF992256D210FA8FFFC7" box="[1287,1341,873,897]" pageId="62" pageNumber="61">Isalo</collectingCounty>
, 
<collectingMunicipality id="6BC9ACE5FF8FFF992203D210FCDDFFE7" pageId="62" pageNumber="61">Ananalava forest</collectingMunicipality>
margin, 
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(
<location id="8ECD6044FF8FFF99231FD2F0FB07FFE7" LSID="urn:lsid:plazi:treatment:03BB8789FF8FFF992463D371FB2EFF87:8ECD6044FF8FFF99231FD2F0FB07FFE7" box="[1102,1205,905,929]" county="Isalo" latitude="-22.5838" longLatPrecision="1" longitude="45.127865" municipality="Ananalava forest" name="Mandabe" pageId="62" pageNumber="61" stateProvince="Fianarantsoa">Mandabe</location>
) vill. env., 
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a. s. l.
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, 
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(
<collectionCode id="ED03AE5AFF8FFF992319D2D0FB23FF87" box="[1096,1169,937,961]" pageId="62" pageNumber="61">FKCP</collectionCode>
)
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.
</paragraph>
</subSubSection>
</treatment>
</document>