Descriptions of New Species of the Diverse and Endemic Land Snail Amplirhagada Iredale, 1933 from Rainforest Patches across the Kimberley, Western Australia (Pulmonata: Camaenidae) Author Köhler, Frank text Records of the Australian Museum 2011 2011-11-30 63 2 167 202 http://dx.doi.org/10.3853/j.0067-1975.63.2011.1581 journal article 10.3853/j.0067-1975.63.2011.1581 2201-4349 5239547 1BCD4085-D2B9-400D-B504-8C85C30303D6 Amplirhagada moraniana n.sp. Type locality . Western Australia , NW Kimberley , 25.6 km SSW of Mitchell River Homestead , N bank of Moran River , 15°20'30"S 125°42'05"E (RFS-18-2: coll. V . Kessner, 18 Jun 1987 ) ( Fig. 1 ) . Type material . Holotype WAM S34713 (preserved specimen) (Pl. 1.4; Table 1 ) . Paratypes WAM S34714 (2 preserved specimens) , FMNH 220725 (3 preserved specimens) , FMNH 220724 (20 dried shells) , WAM S34715 (15 dried shells) , AM C.472922 (6 dried shells). Etymology . Named after the Moran River. Description Shell (Pl. 1.4; Fig. 11 A–E ). Broadly conical with low to moderately elevated spire; thin, translucent. Periphery well rounded to slightly angulate; upper and basal sectors of whorls rounded. Umbilicus 70–90% concealed by columellar reflection. Background and ventral colour pale brownish horn; peripheral and sub-sutural bands usually absent or diffuse, visible on last whorl only; outer and inner lip whitish. Protoconch ~ 2.5 mm in diameter, comprising about 1.5 whorls, with radially elongated pustulations. Teleoconch with regular axial lirae, evenly distributed across shell surface. Angle of aperture about 45 degrees; outer lip thin, well rounded, slightly expanded, slightly reflected; basal node absent or weak. Parietal wall of inner lip inconspicuous. Average shell size 10.9±1.5 × 15.9± 1.5 mm ( Table 1 ). Radular and jaw morphology ( Fig. 1 F–J ). Tooth formula C + 12–13 + 3–4 + 16–18, with 120 rows of teeth (n = 1). Jaw with nine plates. Genital morphology ( Figs 12–13 ). Penis straight, tubular, longer than anterior part of oviduct. Penial sheath very delicate. Length of penial retractor muscle very short, equivalent to about 1/10 of length of penial complex. Penial verge very long, extending about ¾ of length of penial chamber, slender with pointed tip. Inner penial wall almost entirely covered by reloped, rhomboid pustules that are arranged in honey-comb pattern. No main stimulatory pilaster differentiated; three to four longitudinal pilasters formed by fused, more elevated pustules are present, extending most of inner penial chamber, giving rise to corrugated longitudinal pilasters comprising distal end of inner penial wall. Vas deferens very thick, not undulating, entering penial sheath within proximal third of penial complex. Vagina long, tubular; inner vaginal wall and inner wall of bursa copulatrix with continuous, well-developed, smooth longitudinal pilasters. Bursa copulatrix slightly extending base of spermoviduct. Free oviduct comprising less than half of length of anterior part of oviduct. Spermoviduct about as long as anterior part of oviduct. Figure 12. Genitalia of Amplirhagada moraniana n.sp. (WAM S34714 ). For labelling of structures see Fig. 3. Scale bar = 5 mm. Aestivation strategy : Free sealer. Remarks . Anatomical description based on dissection of one specimen . Listed by Solem (1991) as “ Amplirhagada NSP 20”. One of the smallest species with respect to shell height and diameter; a similar shell with respect to size and shape is only found in A. angustocauda described further below. Elongated shape of penis and markedly elongated penial verge (almost as long as penial chamber), corrugated penial wall sculpture, and very thick vas deferens is combination of features typical only for this species.