Cochlostoma Jan, 1830 revised: an overview of the subgenus Turritus Westerlund, 1883 and its species (Caenogastropoda, Cochlostomatidae) Author Zallot, Enrico 365E17AD-6938-4364-A526-F1BDA2663E10 Per Ligont 1, 33070 Budoia, Italy. ezallot@gmail.com Author Kamchev, Panche 9FA2BE01-3355-4340-B57C-9E2841D5E4C2 Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, the Netherlands. panekamchev@gmail.com Author Schilthuizen, Menno 683D0AB7-CDD8-4FAA-94B6-436F3BFB8873 Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, the Netherlands. & Taxon Expeditions, Rembrandtstraat 20, 2311 VW Leiden, the Netherlands. menno.schilthuizen@naturalis.nl Author Fehér, Zoltán E801EC76-8B1E-450B-993E-BBBE57C00EA9 WWF Hungary, Álmos vezér útja 69 / A, 1141 Budapest, Hungary. feher.zoltan.nhmus@gmail.com Author Mattia, Willy De C74A049D-9D7B-4A1F-878B-5BDFD104219F Central Research Laboratories, Burgring 7, 1010, Vienna, Austria; Department of Evolutionary Biology, University of Vienna, Djerassiplatz 1, 1030, Vienna, Austria. wdemattia@gmail.com Author Gittenberger, Edmund D786C279-FC92-4D08-AF16-F79A9705E0AE Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, the Netherlands. egittenberger@yahoo.com text European Journal of Taxonomy 2024 2024-03-21 927 1 163 https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2475/11013 journal article 10.5852/ejt.2024.927.2475 2118-9773 10853688 B6E43365-FACA-49BA-8CCD-77E4BF8F0016 Cochlostoma ( Turritus ) patulum ( Draparnaud, 1801 ) Figs 14 (red dots), 15E, 19–20 Cyclostoma patulum Draparnaud, 1801: 39 . Lectotype (here designated) FRANCE • probably ♀ ; Hérault , Montpellier ; “ Cyclostoma patulum Coll. Draparnaud 1820 XXVI /13” and “ Cochlostoma patulum (Drap.) Typus von Draparnaud!”; NHMW14710 /1. Fig. 18. Syntypes of Cochlostoma ( T. ) zawinkanum ( Wagner, 1906 ) . A . ♀ (NHMW-K51883/1). B . ♂ (NHMW-K51883/2). Other specimens FRANCE • 2- Cirque d’Archiane ; 44.7590° N , 5.4186° E ; 1975; RMNH leg.; RMNH117454 • 3- Cassis ; 43.2157° N , 5.5389° E ; 2009; Pocaterra leg.; EZ0457 • 4- Cirque de Navacelles ; 43.8945° N , 3.5115° E ; 1978; de Winter leg.; A. de Winter-0882 • 5- Gorges de Galamus ; 42.8402° N , 2.4794° E ; 1987; de Winter leg.; A. de Winter 5601 . Type locality FRANCE • Hérault, Montpellier (“H.a.F.M.” in the original description =France meridionale. Wagner (1897) wrote: “Ich beurtheile diese Art nach Originalexemplaren Draparnaud’s von Montpellier in Südfrankreich, welche sich im k. k. naturhistorischen Hofmuseum in Wien befinden”.) Fig. 19. Lectotype (here designated) of Cochlostoma ( T. ) patulum ( Draparnaud, 1801 ) , 1- Monpellier, FR ((NHMW-14710). Fig. 20. Cochlostoma ( T. ) patulum ( Draparnaud, 1801 ) . A–B . 2- Cirque d’Archiane, FR (RMNH- 117454). C . 3- Cassis, FR (EZ-0457. D . 6- Cirque de Navacelles, FR (AJW-0882). Description SHELL . Closely spaced riblets on last part of protoconch. Teleoconch spotless, with strong ribs, slightly weaker and irregularly spaced approaching aperture. Strong and developed lip with columellar lobe abruptly inwardly curved to cover umbilicus. MEASUREMENTS . 4 ♀♀ : whorls= 7.2–8.1, H = 7.4–8.5 mm , H/W= 2.41–2.7, roundness= 0.12–0.13, ribs incl.= 61–64°, apert. incl.= 12–15°, ribs/mm 1 st wh.= 10–15, ribs/mm 4 th wh.= 11–13. FEMALE GENITAL ORGANS . As in C. ( T. ) stussineri . Remarks Cochlostoma ( T. ) patulum from Cirque D’Archiane and from Luberon (GENBANK, GQ370416) do not cluster in the 16S analysis as do conspecific samples belonging to other species. The p-distance between them is 1.5%, higher than the one (1.2–1.3%) found between patulum and stussineri . For this reason, it seems appropriate to better define the type locality, here restricted to Montpellier ( France ). Maybe ‘ancient’ DNA could be amplified from the lectotype , or a specimen of the same sample, to stabilize the taxonomy of the French Turritus .