Morphological investigation of genital organs and first insights into the phylogeny of the genus Siciliaria Vest, 1867 as a basis for a taxonomic revision (Mollusca, Gastropoda, Clausiliidae) Author De Mattia, Willy https://orcid.org/0000-0002-0056-467X Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria willy.demattia@icgeb.org Author Reier, Susanne Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria Author Haring, Elisabeth Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria text ZooKeys 2021 2021-12-14 1077 1 175 http://dx.doi.org/10.3897/zookeys.1077.67081 journal article http://dx.doi.org/10.3897/zookeys.1077.67081 1313-2970-1077-1 C28AD65A76F242CFBED7DFB3702CABCE 734088641608531C8E2CC69397B000ED Stigmatica stigmatica stigmatica ( Rossmaessler , 1836) Figs 59.1-59.6, 61.1-61.3 Distribution. The nominate subspecies is distributed from southern Dalmatia (from the Neretva River southward), Herzegovina, Montenegro to northern Albania. Specimens examined. Croatia , Dubrovnik , Sustjepan , Reka Dubrovacka . 5 m asl , 42°40'12.26"N , 18° 05'38.24"E , W. De Mattia and J. Macor leg. and det., 2 dissected spm ; Montenegro , Rijeka Rezevici , on the Budva-Petrovac road, 100 m asl , 42°13'33.84"N , 18°54'40.70"E , Deli , Eross , Feher leg. W. De Mattia det., 2 dissected spm .; Montenegro , Orjen Mts , Podi and Kameno , Herceg Novi-Trebinije Road. 325 m asl , 42°28'8.05"N , 18°32'29.17"E Deli , Eross , Feher leg. W. De Mattia det., 2 dissected spm . External morphology of the genital organs (Figs 59 .1, 59.3, 59.5). The V is much shorter than the FO. The FDBC is longer than the SDBC+BC. The BC+SDBC is club-like in shape, with a more or less clear distinction between the SDBC and the BC. The D is extremely longer than the SDBC+BC but remarkably thinner. The V is very short but wide in diameter. The PC is ~ 4-5 x longer than the V. The P is cylindrical or slightly swollen. The transition between P and EP is clearly visible. The PR is short but very robust. The E is usually as long as the P and thinner in diameter. Internal morphology of the genital organs (Figs 59 .2, 59.4, 59.6). The V is smooth. The P shows a pattern of irregularly fringed or smooth pleats, sometimes forming and irregular net-like pattern, usually disappearing toward the A that is smooth. The pseudopapilla small and roundish. It originates from the E wall, which is connected with the ELP. The epiphallar formula is ER(PP+ELP). The E shows two fringed ELP, sometimes remarkably irregular and connected with small fleshy transversal bridges. They proximally fade before the VD.