The taxonomy of the American Ariadna Audouin (Araneae: Synspermiata: Segestriidae)
Author
Giroti, André Marsola
Author
Brescovit, Antonio Domingos
text
Zootaxa
2018
2018-03-26
4400
1
1
114
journal article
30419
10.11646/zootaxa.4400.1.1
2b7bf048-65d5-4814-bbdb-59d53d15b00e
1175-5326
1207641
59CC08B8-5B13-42EC-943E-C930D7D0DD7F
Ariadna boliviana
Simon, 1907
Figures 3‒5
,
7‒8
,
10
,
36‒37
,
71
Ariadna boliviana
Simon, 1907
: 262
(♀ lectotype and ♂ lectoparatype from Espíritu Santo, [16°55'S; 67°48'W], BoliVia, Garlepp leg., deposited in MNHNP-AR 14382 and 14383, respectively, designated by
Beatty (1970)
, examined).—
Petrunkevitch (1911: 130)
.—
Beatty (1970: 477, figs 4, 6, 14, 51‒52, 54‒55)
.—
World Spider Catalog (2018)
.
Ariadna dubia
Mello-Leitão, 1917
: 82
(♀ holotype from Pinheiro (actually Piraí, Pinheiral [22°30'46"S, 44°00'03"W], Rio de Janeiro, Brazil, E. Brito leg., should be deposited in MNRJ 1452, lost).—
Beatty (1970: 483, as
nomem dubium
)
.—
Moreira
et al.
(2010
: 61).—
NEW SYNONYMY
.
Synonymy.
Mello-Leitão (1917)
described
A. dubia
based in a dried and pinned specimen, with a deformed and seared abdomen.
Beatty (1970)
, due to the unavailability of the
holotype
and the poor description of Mello-Leitão, treated it as
nomem dubium
. There are two species registered to the type-locality of
A. dubia
:
Ariadna boliviana
Simon, 1907
and
Ariadna obscura
(
Blackwall, 1858
)
. According to the macrosetae pattern presented by
Mello-Leitão (1917)
in
A. dubia
's original description, the tibiae I‒II presents 2-2-2-2 pattern (vp1-0-1-0-1-0-1, vr1-0-1-0-1-0-1) and metatarsi I and II with seven pairs of macrosetae. This macrosetae pattern is consistent with the one from
A. boliviana
, and differs from
A. obscura
. Based in the characters presented by
Mello-Leitão (1917)
,
A. dubia
is considered junior-synonymy of
A. boliviana
.
Additional material examined:
SURINAME
:
Sipaliwini District
:
Upper Kaburi Creek
[
03°46'37"N
;
56°01'40"W
],
1♀
,
IX.1977
,
J.E.D. Milner
leg. (
NHM
)
;
BRAZIL
:
Pará
State:
Parauapebas, Gruta NIN8 N1-174 (
6°01'28.5"S
;
50°17'55.0"W
),
4♀
,
17.VII‒4.VIII.2014
,
Carste
et al.
leg. (
IBSP 193496‒193499
)
;
Gruta
NIN8 N1- 174 (
6°01'29.1"S
;
50°17'54.5"W
),
2♀
,
7‒28.I.2015
,
Carste
et al.
leg. (
IBSP 193502
; 193503);
Gruta
NIN8 N1- 0 223 (
6°01'39.8"S
;
50°16'33.2"W
),
1♀
,
4.IX‒6.X.2014
,
Carste
et al.
leg. (
IBSP 193501
)
; Parauapebas, Gruta GEM-1786 (6°06'19"S; 50°08'21"W), 1♀1imm.,
27.VIII.2010
, R.
A. Zampaulo
leg. (
ISLA
);
Gruta
GEM-1623 (
6°07'06"S
;
50°07'56"W
), 1imm.,
16.III.2011
, M
.
P. Oliveira
leg. (
ISLA
);
Gruta
GEM-1764 (
6°07'58"S
;
50°08'14"W
),
1♀
,
25.X.2010
, M
.
P. Oliveira
leg. (
ISLA
);
Gruta
GEM-1715 (
6°07'59"S
;
50°08'06"W
), 1imm.,
4.V.2011
, M
.
P. Oliveira
leg. (
ISLA
);
Gruta
GEM-1739,
1♀
1imm.,
5.V.2011
, M
.
P. Oliveira
leg. (
ISLA
);
Gruta
GEM-1739,
1♀
1imm.,
31.X.2010
, M
.
P. Oliveira
leg. (
ISLA
);
Gruta
GEM-1799,
1♀
1imm.,
5.V.2011
, M
.
P. Oliveira
leg. (
ISLA
);
2♀
2imm.,
31.X.2010
, R
.
A. Zampaulo
leg. (
ISLA
);
Carajás
,
Floresta Nacional de Carajás
,
Gruta
N4E32 (
6°02'25"S
;
50°09'43"W
),
1♀
,
8‒12.II.2007
,
R. Andrade
et al.
leg. (
IBSP 97751
)
;
Gruta
N4WS80 (
6°04'34.4"S
;
50°11'27.9"W
),
1♀
,
18.XI‒1.XII.2010
(
IBSP 174229
)
;
Gruta
N5S02 (
6°05'33"S
;
50°07'35"W
),
1♀
,
3‒13.V.2005
,
Andrade
&
Arnoni
leg. (
IBSP 55777
)
;
Gruta
N5S-12 (
06°06'11''S
;
50°07'31''W
),
1♀
,
14‒23.X.2009
,
R. Andrade
&
I. Cizauskas
leg. (
IBSP 174230
)
;
Gruta
N5S53/53 (
6°06'28.3"S
;
50°08'01.5"W
),
1♀
,
25.VIII‒3.IX.2009
,
R. Andrade
et al.
leg. (
IBSP 193494
)
;
Gruta
NIN8 N8-031 (
6°09'54.0"S
;
50°09'30.0"W
),
1♀
,
17.VII‒4.VIII.2014
,
Carste
et al.
leg. (
IBSP 193500
)
;
Gruta
N4E31 (
6°05'33"S
;
50°07'35"W
),
1♀
(
IBSP 97752
)
;
Gruta
CRIS-10 (
6°26'56.4"S
;
49°41'08.6"W
),
1♀
,
29.VII‒6.VIII.2008
,
R. Andrade
et al.
leg. (
IBSP 174570
)
;
Novo Progresso
,
Serra do Cachimbo
,
Campo de Provas Brigadeiro Velloso
(09°21'89"S;
55°02'01"W
),
1♀
,
11.IX.2003
,
D.R. Santos-Souza
leg. (
MPEG 6243
)
; without specific data, 1♀, Mth. (MNHNP-AR);
Maranhão
State:
Caxias, Reserva Ecológica Inhamum (
04°53'56"S
;
43°26'10"W
),
2♂
,
16‒19.I.2006
, G. A
. Cunha leg. (IBSP
122448
); 1♂,
12‒15.VII.2006
, G.A. Cunha leg. (IBSP
122449
); 1♂, 2006, F. Limeira leg. (IBSP
122450
);
Piauí
State:
Teresina
[
05°05'20"S
;
42°48'07"W
],
Sítio do Carlos
Brasil
,
1♀
,
15.V.2004
,
L.S. Carvalho
leg. (
MPEG 5320
)
;
Areias
,
Parque Estadual Mata do Pau Ferro
(
06°58'38,6"S
;
35°42'25"W
),
1♂
,
26‒30.
IX.2011
,
E. Araújo
&
A. Medeiros
leg. (CHNUFPI 0285)
;
Pernambuco
State:
Tapera
[
09°25'38"S
;
40°44'15"W
],
1♀
,
1.III.1972
,
P.F.L. Duarte
leg. (
MCN
25804)
;
Sergipe
State:
Itabaiana
,
Parque Nacional da Serra de Itabaiana
,
161m
(
10°47'02.9''S
;
37°20'09.5''W
),
1♀
,
20.V.2012
,
I.L.F. Magalhães
&
J.L. Chavari
leg. (
IBSP 166856
)
;
Mato Grosso
State:
Cotriguaçu, Fazenda São Nicolau (
09°50'24"S
;
58°14'54"W
),
1♂
1♀
,
2‒14.X.2010
,
A.J. Santos
leg. (
UFMG 7743
; 7744)
;
Miracema do Norte
[
09°34'01"S
;
48°23'31"W
],
1♀
1imm.,
17‒25.IV.2005
,
I. Knysak
&
R. Martins
leg. (
IBSP 124516
)
; Porto Nacional, Capivara [10°09'44"S; 48°31'07"W], 1♀,
21.IX.2000
, I. Knysak & R. Martins leg. (IBSP 124057);
Porto Nacional
[
10°42'28"S
;
48°25'01"W
], Fazenda Sandoval, Cerrado,
1♀
,
19.IX.2002
,
R. Martins
&
I. Knysak
leg. (
IBSP 123806
)
;
Palmas
[
10°11'04"S
;
48°20'01"W
],
1♀
,
30.IX.2001
,
E. Butantan
leg. (
IBSP 37797
)
; Querência, Fazenda Tanque (13°04'43''S; 52°23'26''W), 1♀,
I.2014
, R.B. Almeida leg. (IBSP 169568);
Sapezal
[
13°32'33"S
;
58°48'51"W
],
Utiariti
,
1♀
, 1961,
K. Lenko
leg. (
MZSP 3583
)
;
Vale de São Domingos
/
Pontes de Lacerda
,
Usina Hidrelétrica de Guaporé
[
15°07'S
;
58°58'W
],
1♀
,
X.1999
,
E. Resgate
leg. (
IBSP 41650
)
;
Cuiabá, Gruta Kiogo Brado (
15°37'1.94"S
;
55°30'16.61"W
),
1♀
2imm.,
27 October 2006
,
R.L. Ferreira
leg. (
ISLA
)
;
Tocantins
State:
Palmas
,
Gruta da Boa Esperança
(
10°22'55.3"S
;
48°03'35.2"W
),
2♀
,
2.XII.2004
, R
.
L. Ferreira
leg. (
ISLA
);
Goiás
State:
São Domingos
[
13°23'52"S
;
46°19'04"W
],
Parque Estadual de Terra Ronca
,
Cavernas Passa Três
,
1♀
,
3‒5.IX.2000
, F
.
P. Franco
leg. (
IBSP 26082
);
Minaçu
,
Usina Hidrelétrica Serra da Mesa
[
13°50'01"S
;
48°18'10"W
],
2♂
3imm.,
18‒30.XI.1996
, A
.
B. Bonaldo
&
L. Moura
leg. (
MCN
27861);
São Jorge
,
Parque Nacional da Chapada
dos
Veadeiros
[
14°10'S
;
47°30'W
],
1♀
,
30.X.2009
, P
.C. Motta leg. (DZUB 5379);
Cocalzinho de
Goiás
[
15°47'38"S
;
48°46'33"W
],
RPPN
,
1♂
1imm., 2008,
I.P. Rocha
leg. (
DZUB 5965
)
;
Morrinhos
[
17°43'55"S
;
49°06'03"W
],
Fazenda Recanto do Jatobá
,
2♂
, 2009,
W.C.R. Pinto
leg. (
IBSP 152775
; 152777);
Jataí
[
17°52'51"S
;
51°42'50"W
],
2♀
,
20.IX.2005
, P
.
C. Motta
leg. (
DZUB 3880
); Catalão, Barragem para
Aproveitamento Hidrelétrico Serra do Facão
(
18°03'20,07"S
;
47°37'55,63"W
),
1♀
,
VII.2010
, R
.
B. Martines
&
R.M.C. Silveira
leg. (
UFMG 4526
);
Catalão
[
18°10'12"S
;
47°56'31"W
],
1♂
,
20.X.2001
, P
.
C. Motta
,
F.S.P. Godói
&
D. Diniz
leg. (
DZUB 2530
);
Bahia
State:
Porto Seguro
,
Parque Nacional do Monte Pascoal
(
16°47'31"S
;
39°28'30"W
),
1♀
,
21‒23.IV.1998
, A
.
D.
Brescovit
et al.
leg. (
IBSP 18438
);
Mato Grosso
do Sul
State:
Corumbá
[
19°00'32"S
;
57°39'10"W
],
Serra do Urucum
,
1♀
,
27.XI.1960
, K
.
Lenko
&
C. Carvalho
leg. (
MZSP 11564
);
Paranaíba
[
19°40'37"S
;
51°11'27"W
],
1♂
,
XI.1983
, R
.R. da Silva leg. (IBSP 3768); 1♀,
I.1983
(IBSP 4141); 1♀,
7.VI.1983
(IBSP 14084);
Dois Irmãos do Buriti
[
20°27'S
;
55°30'W
],
Piraputanga
,
1♀
,
31.VI‒6.VII.1999
,
A. D. Brescovit
et al.
leg. (
IBSP 98896
)
;
Três Lagoas
[
20°45'04"S
;
51°40'42"W
],
1♀
,
VI.1964
,
C.D.
Z. leg. (
MZSP 3616
)
; 1♀,
X.1964
, Exp. O.Z. leg. (MZSP 3766);
Três Lagoas
,
Horto Barra da Moeda
(
20°58'S
;
51°47'W
),
1♂
,
XI.2007
,
M. Uehara-Prado
leg. (
UFMG 3202
)
;
Santa Rita do Rio Pardo
[
21°18'10"S
;
52°49'51"W
],
1♀
,
23.IV.2001
,
R. Bertani
&
E.K. Kashimata
leg. (
IBSP 39480
)
;
Minas Gerais
State:
Januária
,
Parque Nacional do Peruaçu
,
550m
(
15°08'45''S
;
44°14'28.6''W
),
1♀
,
15.IV.2015
, L
.
S. Carvalho
leg. (CHNUFPI 1516);
Itacarambi
,
Distrito de Fabião
II (
15°10'19"S
;
44°10'49"W
),
1♂
,
17‒22.X.2010
,
G.F.B.P. Ferreira
et al.
leg. (
UFMG 9357
)
;
Águas Vermelhas
[
15°44'49"S
;
41°27'36"W
],
1♂
,
III.1985
,
V. Palmeira
leg. (
MNRJ 6
415)
;
Diamantina
,
Minas de Serrinha
[
18°14'56"S
;
43°36'00"W
],
1♂
, 1945,
E. Cohn
leg. (
AMNH
)
;
Morro do Pilar
,
Lageado
(
19°13'47,28''S
;
43°22'59,76''W
),
7♀
1imm.,
13‒14.XI.2013
,
P.H. Martins
et al.
leg. (
UFMG 14361
)
;
Sete Lagoas
,
Gruta do Chisé
[
19°27'57"S
;
44°14'49"W
],
1♀
,
25.II.2009
(
IBSP 149911
)
;
Sete Lagoas
,
Entorno da Gruta da Lapinha
(
19°33'S
;
43°57'W
),
1♀
,
23.VI.2001
,
R.L.C. Baptista
leg. (
MNRJ 6186
)
; Matozinhos, Gruta MOC113 (19°33'19.3"S; 50°01'15.0"W), 1♀,
1‒8.VIII.2011
, F.
Franco
leg. (
IBSP 19
3495);
Timóteo
,
Parque Estadual do Rio Doce
(
19°39'28.3"S
;
42°34'22.5"W
),
2♂
,
19.XI.2010
, B
.
T. Faleiro
leg. (
UFMG 10501
);
Timóteo
,
Parque Estadual do Rio Doce
(
19°39'30,1"S
;
42°34'25,2"W
),
1♂
,
19.XI.2010
, B
.
T. Faleiro
leg. (
UFMG 10488
);
Parque Estadual do Rio Doce
(
19°39'30,6"S
;
42°34'28,2"W
),
1♂
,
19.XI.2010
, B
.
T. Faleiro
leg. (
UFMG 10476
);
Parque Estadual do Rio Doce
(
19°39'30,7"S
;
42°34'32,2"W
),
2♂
,
17.XII.2010
, B
.
T. Faleiro
leg. (
UFMG 10457
);
Marliéria
,
Parque Estadual do Rio Doce
(
19°42'58,62''S
;
42°44'01,69''W
),
2♂
,
2‒9 November 2000
, D
.P. Araújo leg. (UFMG 14338);
Caeté
,
Serra da Piedade
[
19°49'23.3"S
;
43°41'02.4"W
],
1♀
,
16.XII.2012
,
Oliveira
leg. (
IBSP 209936
)
;
Santa Bárbara
[
19°57'32"S
;
43°24'54"W
],
Reserva Florestal Peti
,
1♂
1♀
,
14.VIII.1986
,
R.L.C. Baptista
leg. (
MNRJ 6407
)
;
Catas Altas
[
20°04'30"S
;
43°24'28"W
],
Serra do Caraça
,
1♀
,
12‒23.XI.1961
,
U. Martins-Lenko
&
R. Kloss
leg. (
MZSP 6687
)
;
Catas Altas
,
RPPN
Parque Natural do Caraça
(
20°05'51"S
;
43°29'18"W
),
1♀
, 2011 (
UFMG 10368
)
;
Mariana
,
Mina da Alegria
[
20°10'31.3"S
;
43°29'37.4"W
],
Caverna
0 2,
1♀
,
16‒17.IV.2009
,
R. Bessi
et al.
leg. (
IBSP 149763
)
;
Ouro Preto
(
20°13'S
;
43°30'W
),
1♀
,
1.III.2015
,
H.C. Costa
leg. (CHNUFPI 1570)
;
Espírito Santo
State:
Linhares
,
Floresta Nacional de Goytacazes
(
19°26'07"S
;
40°04'32"W
),
1♀
,
26.X.2005
,
T. Souza
et al.
leg. (
IBSP 134971
)
; Santa Teresa, Reserva Biológica Augusto Ruschi [19°54'20"S; 40°33'44''W], 1♀,
VI‒VIII.2006
, T. Souza, T. Barnabé & E. Soveiro leg. (IBSP
121566
);
São Paulo
State:
Luís Antônio
,
Estação Ecológica de Jataí
[
21°37'03''S
;
47°45'55''W
],
5♂
1♀
,
3‒7.X.2009
,
A.G. Cristovão
leg. (
IBSP 166947‒166951
)
; 1♀,
6‒12.XI.2009
, A.G. Cristovão leg. (IBSP 166946);
Tabatinga
[
21°47'05"S
;
48°38'09"W
],
Mata Sagrada
,
1♀
,
27.VIII.1965
,
K. Lenko
leg. (
MZSP 1
1565)
; Araraquara, Jardin Nova América [21°47'38"S; 48°10'33"W], 1♀,
XII.2010
, S.C.F.S.A. leg. (IBSP 209935);
Assis
,
Estação Ecológica de Assis
(
22°34'S
;
50°24'W
),
1♂
1♀
,
25‒30.XI.2002
,
Exp
.
Biota
leg. (
IBSP 98907
; 98908);
Assis
[
22°39'43"S
;
50°24'43"W
],
3♂
,
F.T.S. Morimoto
leg. (
IBSP 166852‒166854
)
; Rosana, Usina Hidrelétrica Engenheiro Sérgio Motta [22°34'48"S; 53°03'32"W], 1♀,
I‒II.2000
, E. Butantan leg. (IBSP 29705);
Bernardino de Campos
[
23°00'46"S
;
49°28'26"W
],
1♀
,
19.XI.2003
,
A.C.B. Silveira
leg. (
IBSP 40946
)
;
Ubatuba
,
Parque Estadual da Ilha Anchieta
(23°31'‒
23°34'S
; 45°02'‒
45°05'W
),
1♂
,
23‒30.VII.2001
,
Exp. Biota
leg. (
IBSP 66834
)
;
São Paulo
[
23°32'52"S
;
46°38'09"W
],
1♀
,
X.1991
,
J.C. Ferreira
leg. (
IBSP 5822
)
;
Barueri
[
23°30'39"S
;
46°52'33"W
],
1♀
,
9.I.1966
,
K. Lenko
leg. (
MZSP 5598
)
; 1♀,
3.X.1960
, K. Lenko leg. (MZSP 8387); Registro, Porto Cabral [24°29'16"S; 47°50'38"W], 2♂, L. Travessos F° leg. (MZSP 77; 8591);
Rio de Janeiro
State:
Campos dos Goytacazes
,
Morro Itaoca
,
Aprovisionamento
(
21°45'14"S
;
41°19'26"W
),
1♀
,
22.I.2010
,
M. Marques
leg. (
IBSP 166857
)
;
Paraíba
do Sul
,
Fazenda Maravilha
(
22°12'S
;
43°16'W
),
1♀
,
16‒18.VIII.2001
,
Exp. Biota
leg. (
IBSP 98888
)
;
Macaé
,
Ilha de Santana
[
22°24'S
;
41°42'W
],
1♂
,
19‒20.X.1986
,
R.L.C. Baptista
leg. (
MNRJ 6406
)
;
Pinheiral
,
Fazenda
Santa Helena
[
22°31'S
;
43°58'W
],
3♂
1♀
,
5‒11.XI.1999
,
A.D. Brescovit
et al.
leg. (
IBSP 98884‒98887
)
;
Rio de Janeiro
,
Floresta da Tijuca
,
Guanabara
[
22°57'47"S
;
43°14'40"W
],
3♂
,
VI.1966
,
Alvarenga
leg. (
MZSP 20137
; 20139; 20167)
;
Paraná
State:
Diamante do
Paraná
[
22°37'06"S
;
52°51'23"W
],
Estação Ecológica do Caiuá
,
1♀
, 2012,
D. Araujo
et al.
leg. (
IBSP 166855
)
;
Pinhais
[
22°26'41"S
;
49°11'33"W
],
Serra da Farinha Seca
,
1♀
,
15‒20.IX.1995
,
E.L. Aracnologia
leg. (
MCTP 7649
)
;
Cambará
[
23°02'45''S
;
50°04'26''W
],
1♀
,
15‒17.VI.1937
,
H.E. Hinton
leg. (
NHM
1941.1.7.63)
;
BOLIVIA
:
Santa
Cruz Department
:
Buena Vista
,
344m
(
17°29'33,6"S
;
63°39'29,9"W
),
1♂
,
2.XII.2010
, R
.
Pintoda-Rocha
et al.
leg. (
MZSP 68119
);
PARAGUAI
:
1♀
,
1891‒1893
,
Dr
. Bohls leg. (NHM 1906.3.24.3-1500 (hart)).
Diagnosis.
Ariadna boliviana
is similar to
A. isthmica
,
A. ipojuca
n. sp.
and
A. gaucha
n. sp.
due to the morphology of male palp (
Figs 37A‒B
,
44A‒B
,
58G‒H
,
61A‒B
), metatarsi IV (
Figs 36F
,
43F
,
58F
,
60F
), and leg macrosetae pattern of females, but differ by the presence of males with sinuous metatarsi I, with a marked bend in the proximal region (
Fig. 36C‒E
); females differ from these species, except
A. gaucha
n. sp.
, by the presence of a ventral lobe longer than the dorsal one; it differs from
A. gaucha
n. sp.
by the less sinuous ventral lobe (
Figs 10A‒C
,
37C‒D
).
Description. Male (IBSP 152775).
Carapace and chelicerae brownish orange (
Fig. 36A
). Endites orange; labium dark orange. Sternum orange with darker margins, and three pairs of dark marginal areas, the hind one fainted (
Fig. 36B
). Palps light orange (
Fig. 37A‒B
). Legs brownish orange, pairs I‒II darker, with the femur, patella and tibia marbled with dark pigment, and dark diffuse rings in the distal region of metatarsus (
Fig. 36C‒F
). Abdomen grey, lighter ventrally (
Fig. 36A‒B
). Total length 8.64; carapace 4.12 long, 2.8 wide. Palps: tibia incrassated, with the dorso-ventral diameter shorter than the bulb; bulb sub-rectangular, mid-portion shorter than the embolus and directed backwards; mid-portion/embolus transition conspicuous; embolus long and with a tubular tip (
Figs 4A‒C
,
37A‒B
). Legs: robust and coated with setae (
Fig. 36C‒F
); metatarsi I with short macrosetae, with similar size (
Fig. 36C‒E
); metatarsi IV sinuous, in lateral view (
Fig. 36F
); tarsal brush of putative chemosensory setae present in legs II‒IV, extending to the median region of the metatarsi (
Fig. 36F
). Leg formula: I-II-IV-III. Leg measurements: I, femur 3.92, patella 1.6, tibia 3.56, metatarsus 3.84, tarsus 0.84, total 13.76; II, 3.76, 1.6, 3.44, 3.6, 0.84, 13.24; III, 2.68, 1.28, 2.04, 2.24, 0.76, 9.0; IV, 3.16, 1.48, 2.6, 2.32, 0.8, 10.36. Macrosetae: I, femur d0-0-1- 0-1-0-1-0-2, p0-0-0-0-0-1-1-0; patella p0-0-1/0; tibia d0-0-0-1/0-0-1-0, r1-1-0-1-0-1-0, vp1-0-0-0-0-0-1, vr2-0-1- 0-1-0/1-1, r1-1-1-1-1-1-1-1; metatarsus p0-0-1-0-0-0-0-0-0-0, vp0-0-0-1-1-1-1-1-0-0, vr0-0-0-0-1-0-0-1 (
Fig. 36C‒E
); II, femur d0-0-1-0-1-0-1-1-2, p0-0-0-0-0-1-1-0; tibia d0-0-0-0-0-1-0, p1-1-0-1-0-1-0, vp0-0-0-0-1-0-1, vr2-0-1-0-1-0-1, r1-1/0-1-1-2/1-0-1-0; metatarsus p0-0-1-0-0-0-0, vp0-0-1-0-0/1-1-0-1, vr0-1-0-1-1-0-1-1, r0-0-1- 0-0-1-0-0; IV, D1-1-1-0-1-0-0-0, r0-0-0-0-0-0-1-0; IV, metatarsus vp0-0-0-0-0-0-1, vr metatarsal comb with 4 macrosetae.
Female (IBSP 193500).
Carapace and chelicerae dark reddish brown (
Fig. 36G
). Endites brownish orange; labium dark brownish orange. Sternum brownish orange with darker margins, and three pairs of dark marginal areas, the hind one fainted (
Fig. 36H
). Palps brownish orange, darkening gradually in distal direction. Legs brownish orange, pairs I‒II darker and with dark pigment densely distributed in the femur, patella, tibia, and proximal region of the metatarsus, and forming rings in metatarsus and tarsus distal region (
Fig.
36I
‒J
). Abdomen as in male (
Fig. 36G‒H
). Total length 12.4; carapace 5.44 long, 3.6 wide. Carapace with the cephalic region elevated (
Figs 1B
,
36G
). Legs: robust and densely coated with setae, mainly on the ventral region of pairs I‒III; legs I‒II with ventro-prolateral and ventro-retrolateral tibial macrosetae of similar size, and metatarsal macrosetae of different sizes (
Fig. 7A‒C
,
36I
‒J
). Leg formula: I-II-IV-III. Leg measurements: I, femur 4.28, patella 2.12, tibia 3.6, metatarsus 3.44, tarsus 0.88, total 14.32; II, 4.0, 2.04, 3.52, 3.32, 0.84, 13.72; III, 2.96, 1.56, 2.24, 2.24, 0.8, 9.8; IV, 3.76, 1.84, 2.92, 2.44, 0.8, 11.76. Macrosetae: I, femur d0-0-0-0-0-0-2, p0-0-0-0-2-1-0; tibia p0-1-0-1-0-1- 0, vp1-0-1-0-1-0-1, vr1-0-1-0-1-0-1, r0-1-0-1-0-1-0; metatarsus vp1-1-1-1-1-1-1-1-1, vr1-1-1-1/0-1-1-1-1-1 (
Fig.
36I
‒J
); II, femur d0-0-0-0-0-2-0, p0-0-0-0-1-1-0; tibia p0-1-0-1-0-1-0, vp0-0-1(slender)-0-1-0-1, vr1-0-1-0-1-0-1, r0-1-0-1-0-1-0; metatarsus vp1-1-1/0-1-1-1-1-1-1-1, vr1-1-1-0-0-1-1-0-1-1; IV, metatarsus vp0-0-0-0-0-0-1, vr metatarsal comb with 4 macrosetae (
Fig. 7F
). Internal genitalia: anterior receptaculum with glandular ducts plate located in the median region, expanded in its proximal region (
Figs 10A‒C
,
37C‒D
).
Variation.
Male (n=10): Total length 5.2‒8.64; carapace 2.8‒4.12 long, 1.96‒2.8 wide; femur I 2.96‒3.96. Female (n=10): total length 8.4‒12.4; carapace 4.12‒5.76 long, 2.6‒3.8 wide; femur I 2.84‒4.52.
Distribution.
Suriname
,
Bolivia
,
Paraguay
and almost entire Brazilian territory (
Fig. 71
).