<document id="304D0020C2B7A670244BAD9B060F1FE0" ID-DOI="10.11646/phytotaxa.313.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="13700295" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725574047968" checkinUser="felipe" docAuthor="Pederneiras, Leandro Cardoso, Mansano, Vidal De Freitas &amp; Romaniuc-Neto, Sergio" docDate="2017" docId="0391336CFFD9FF97FF1023882D6EF8AA" docLanguage="en" docName="phytotaxa.313.1.1.pdf" docOrigin="Phytotaxa 313 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.313.1.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Ficus aequatorialis Dugand 1942" docType="treatment" docVersion="2" lastPageNumber="3" masterDocId="FFA84B14FFD8FF95FF98253A2E22FFB8" masterDocTitle="Taxonomic synopsis of the Ficus sect. Pharmacosycea (Moraceae) from Colombia" masterLastPageNumber="22" masterPageNumber="1" pageNumber="2" updateTime="1725583850547" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="1581D57EAAA35E1691ABBAA943D4AD1B">Taxonomic synopsis of the Ficus sect. Pharmacosycea (Moraceae) from Colombia</mods:title>
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<mods:namePart id="39B6823735B735C97F946373BC5A587E">Pederneiras, Leandro Cardoso</mods:namePart>
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<mods:namePart id="8C94B87DDF51B6BCCAD5851C2BFDD02F">Mansano, Vidal De Freitas</mods:namePart>
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<mods:namePart id="BE9B7A05D3FAD869977DF40DDE99F78F">Romaniuc-Neto, Sergio</mods:namePart>
<mods:affiliation id="1B6350DBE6F3D56BFD9D2C7A9016463E">Instituto de Botânica, Herbário SP, Caixa Postal 4005, São Paulo-SP, 01061 - 970, Brazil.</mods:affiliation>
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<mods:title id="123230624A7355EFA934B07B9D53E636">Phytotaxa</mods:title>
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<mods:part id="336BA3A245B2C22B45F10773BC70921A">
<mods:date id="23A11BFFB8FF0134E36AAF84E50632C9">2017</mods:date>
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<treatment id="0391336CFFD9FF97FF1023882D6EF8AA" LSID="urn:lsid:plazi:treatment:0391336CFFD9FF97FF1023882D6EF8AA" httpUri="http://treatment.plazi.org/id/0391336CFFD9FF97FF1023882D6EF8AA" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
<subSubSection id="C322D1F1FFD9FF94FF1023882C45F973" box="[136,615,1712,1739]" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="8B87827AFFD9FF94FF1023882C45F973" blockId="1.[136,1452,1712,1810]" box="[136,615,1712,1739]" pageId="1" pageNumber="2">
<heading id="D0CF3516FFD9FF94FF1023882C45F973" box="[136,615,1712,1739]" fontSize="11" level="2" pageId="1" pageNumber="2" reason="4">
<emphasis id="B94C5E68FFD9FF94FF1023882EBCF973" bold="true" box="[136,158,1714,1739]" pageId="1" pageNumber="2">1.</emphasis>
<taxonomicName id="4C38F9F9FFD9FF94FF3C238B2C41F973" ID-CoL="6HVSW" authority="Dugand (1942: 28)" authorityName="Dugand" authorityPageNumber="28" authorityYear="1942" box="[164,611,1712,1739]" class="Magnoliopsida" family="Moraceae" genus="Ficus" kingdom="Plantae" order="Rosales" pageId="1" pageNumber="2" phylum="Tracheophyta" rank="species" species="aequatorialis">
<emphasis id="B94C5E68FFD9FF94FF3C238B2FA1F973" bold="true" box="[164,387,1712,1739]" italics="true" pageId="1" pageNumber="2">Ficus aequatorialis</emphasis>
<bibRefCitation id="EFA9FF8BFFD9FF94FE1223882C41F973" author="Dugand, A." box="[394,611,1714,1739]" pageId="1" pageNumber="2" pagination="25 - 74" refId="ref19960" refString="Dugand, A. (1942) Nuevas especies colombianas de genero Ficus. Caldasia 1: 25 - 74." type="journal article" year="1942">Dugand (1942: 28)</bibRefCitation>
</taxonomicName>
.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C322D1F1FFD9FF94FF1023E32AB2F87F" pageId="1" pageNumber="2" type="materials_examined">
<paragraph id="8B87827AFFD9FF94FF1023E32CF4F8AA" blockId="1.[136,1452,1712,1810]" pageId="1" pageNumber="2">
<materialsCitation id="3B508827FFD9FF94FF1023E32CF0F8AA" collectingDate="1940-12-07" collectionCode="COL, F" collectorName="J. Cuatrecasas" country="Colombia" county="Rio San Miguel" location="Type" municipality="Frontera" pageId="1" pageNumber="2" specimenCount="2" typeStatus="holotype">
<location id="8EE7D4A1FFD9FF94FF1023E32E98F956" LSID="urn:lsid:plazi:treatment:0391336CFFD9FF97FF1023882D6EF8AA:8EE7D4A1FFD9FF94FF1023E32E98F956" box="[136,186,1753,1774]" country="Colombia" county="Rio San Miguel" municipality="Frontera" name="Type" pageId="1" pageNumber="2">Type</location>
:— 
<collectingCountry id="F32FC2EAFFD9FF94FF4E23E32F78F956" box="[214,346,1753,1774]" name="Colombia" pageId="1" pageNumber="2">COLOMBIA</collectingCountry>
. 
<collectingMunicipality id="6BE31800FFD9FF94FEFD23E32F94F956" box="[357,438,1753,1774]" pageId="1" pageNumber="2">Frontera</collectingMunicipality>
colombo-ecuatoriana, 
<collectingCounty id="62E6FAF6FFD9FF94FD0023E32D1AF956" box="[664,824,1753,1774]" pageId="1" pageNumber="2">Río San Miguel</collectingCounty>
, 
<locationDeviation id="9A359B58FFD9FF94FCDC23E32AF2F956" box="[836,1232,1753,1774]" firstLocation="Quebrada del Ocano" pageId="1" pageNumber="2" secondLocation="Rio Teteye">entre Quebrada del Ocano y Río Teteyé</locationDeviation>
, 
<date id="FF86A4BAFFD9FF94FB4523E32BACF956" box="[1245,1422,1753,1774]" pageId="1" pageNumber="2" value="1940-12-07">
<collectingDate id="EFC25D52FFD9FF94FB4523E32BACF956" box="[1245,1422,1753,1774]" pageId="1" pageNumber="2" value="1940-12-07">7 December 1940</collectingDate>
</date>
, 
<emphasis id="B94C5E68FFD9FF94FA0323E32F55F8AA" italics="true" pageId="1" pageNumber="2">
<collectorName id="26CDE7ACFFD9FF94FA0323E32F16F8AA" pageId="1" pageNumber="2">J. Cuatrecasas</collectorName>
10887
</emphasis>
(
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: 
<collectionCode id="ED291ABFFFD9FF94FE7D23C72C3FF8AA" box="[485,541,1789,1810]" country="Colombia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15629" lsid="urn:lsid:biocol.org:col:15629" name="Universidad Nacional de Colombia" pageId="1" pageNumber="2" type="Herbarium">COL</collectionCode>
!; 
<typeStatus id="54833CD8FFD9FF94FDB223C72C5EF8AA" box="[554,636,1789,1810]" pageId="1" pageNumber="2" type="isotype">isotypes</typeStatus>
: 
<collectionCode id="ED291ABFFFD9FF94FD1D23C72CB5F8AA" box="[645,663,1789,1810]" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="1" pageNumber="2" type="Herbarium">F</collectionCode>
! US!)
</materialsCitation>
.
</paragraph>
<paragraph id="8B87827AFFD9FF94FF1022782AB2F87F" blockId="1.[136,1452,1858,2063]" pageId="1" pageNumber="2">
Terrestrial trees, 
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tall. Leaf blade 
<quantity id="4CC02F9FFFD9FF94FDFA22782CF1F8E3" box="[610,723,1858,1883]" metricMagnitude="-1" metricUnit="m" metricValue="3.1" metricValueMax="3.7" metricValueMin="2.5" pageId="1" pageNumber="2" unit="cm" value="31.0" valueMax="37.0" valueMin="25.0">25–37 cm</quantity>
long, 
<quantity id="4CC02F9FFFD9FF94FC8722782D84F8E3" box="[799,934,1858,1883]" metricMagnitude="-1" metricUnit="m" metricValue="1.675" metricValueMax="1.9" metricValueMin="1.45" pageId="1" pageNumber="2" unit="cm" value="16.75" valueMax="19.0" valueMin="14.5">14.5–19 cm</quantity>
wide, elliptic, apex acuminate, base cordate, glabrous adaxially, puberulent to glabrous abaxially, secondary veins 9–16 pairs; petiole 
<quantity id="4CC02F9FFFD9FF94FBC7225C2AF1F8C7" box="[1119,1235,1894,1919]" metricMagnitude="-2" metricUnit="m" metricValue="8.25" metricValueMax="11.0" metricValueMin="5.5" pageId="1" pageNumber="2" unit="cm" value="8.25" valueMax="11.0" valueMin="5.5">5.5–11 cm</quantity>
long, puberulent to glabrous, periderm flaking off; stipules 
<quantity id="4CC02F9FFFD9FF94FDD822B02CB0F81B" box="[576,658,1930,1955]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="1" pageNumber="2" unit="cm" value="7.0" valueMax="8.0" valueMin="6.0">6–8 cm</quantity>
long, subpersistent, glabrous. Syconia in pairs, peduncle sessile; lateral bracts rarely seen; receptacle 
<quantity id="4CC02F9FFFD9FF94FE4B22942C72F87F" box="[467,592,1966,1991]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="2.2" metricValueMin="1.5" pageId="1" pageNumber="2" unit="cm" value="1.85" valueMax="2.2" valueMin="1.5">1.5–2.2 cm</quantity>
in diam. when dry, hispid, apex rounded (
<figureCitation id="13039EFFFFD9FF94FBB122942AA0F87F" box="[1065,1154,1966,1991]" captionStart="FIGURE 1" captionStartId="2.[136,229,1500,1521]" captionTargetBox="[153,1433,190,1470]" captionTargetId="figure-16@2.[151,1436,187,1472]" captionTargetPageId="2" captionText="FIGURE 1. Twigs with leaves and stipules. A. Ficus aequatorialis (Cuatrecasas 11214). B. F. apollinaris (Barbosa 1245). C,D. Ficus carchiana (McPherson 13393)." figureDoi="http://doi.org/10.5281/zenodo.13700297" httpUri="https://zenodo.org/record/13700297/files/figure.png" pageId="1" pageNumber="2">Fig. 1A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C322D1F1FFD9FF97FF2522E82D6EF8AA" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2" type="distribution">
<paragraph id="8B87827AFFD9FF94FF2522E82C39F7B7" blockId="1.[136,1452,1858,2063]" pageId="1" pageNumber="2">
<emphasis id="B94C5E68FFD9FF94FF2522E82CA3F853" bold="true" box="[189,641,2002,2027]" pageId="1" pageNumber="2">
Distribution and habitat in 
<collectingCountry id="F32FC2EAFFD9FF94FD9B22E82C5EF853" box="[515,636,2002,2027]" name="Colombia" pageId="1" pageNumber="2">Colombia</collectingCountry>
:
</emphasis>
— 
<collectingRegion id="49FC4C98FFD9FF94FD0622E82D33F853" box="[670,785,2002,2027]" country="Colombia" name="Putumayo" pageId="1" pageNumber="2">Putumayo</collectingRegion>
, in premontane rainforests on the eastern slope of the Andes at elevations of 
<quantity id="4CC02F9FFFD9FF94FEA222CC2F9BF7B7" box="[314,441,2038,2063]" metricMagnitude="2" metricUnit="m" metricValue="4.25" metricValueMax="6.0" metricValueMin="2.5" pageId="1" pageNumber="2" unit="m" value="425.0" valueMax="600.0" valueMin="250.0">250–600 m</quantity>
(
<figureCitation id="13039EFFFFD9FF94FE5022CC2C2CF7B7" box="[456,526,2038,2063]" captionStart="FIGURE 2" captionStartId="3.[136,229,1465,1486]" captionTargetBox="[158,1430,302,1440]" captionTargetId="figure-63@3.[158,1430,302,1440]" captionTargetPageId="3" captionText="FIGURE 2. Distribution of Ficus aequatoriales, F. apollinaris, F. carchiana, F. crassiuscula, F. crassivenosa, F. dewolfii and F. dulciaria in Colombia based on herbarium specimens." figureDoi="http://doi.org/10.5281/zenodo.13700299" httpUri="https://zenodo.org/record/13700299/files/figure.png" pageId="1" pageNumber="2">Fig. 2</figureCitation>
).
</paragraph>
<caption id="DF47D2F2FFDAFF97FF1020E62F97F9AD" ID-DOI="http://doi.org/10.5281/zenodo.13700297" ID-Zenodo-Dep="13700297" httpUri="https://zenodo.org/record/13700297/files/figure.png" pageId="2" pageNumber="3" startId="2.[136,229,1500,1521]" targetBox="[153,1433,190,1470]" targetPageId="2" targetType="figure">
<paragraph id="8B87827AFFDAFF97FF1020E62F97F9AD" blockId="2.[136,1451,1500,1557]" pageId="2" pageNumber="3">
<emphasis id="B94C5E68FFDAFF97FF1020E62EDDFA49" bold="true" box="[136,255,1500,1521]" pageId="2" pageNumber="3">FIGURE 1.</emphasis>
Twigs with leaves and stipules. A. 
<taxonomicName id="4C38F9F9FFDAFF97FDC320E62D35FA49" authorityName="Dugand" authorityYear="1942" box="[603,791,1500,1521]" class="Magnoliopsida" family="Moraceae" genus="Ficus" kingdom="Plantae" order="Rosales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="aequatorialis">
<emphasis id="B94C5E68FFDAFF97FDC320E62D35FA49" box="[603,791,1500,1521]" italics="true" pageId="2" pageNumber="3">Ficus aequatorialis</emphasis>
</taxonomicName>
(
<emphasis id="B94C5E68FFDAFF97FCBE20E62DFDFA49" box="[806,991,1500,1521]" italics="true" pageId="2" pageNumber="3">Cuatrecasas 11214</emphasis>
). B. 
<taxonomicName id="4C38F9F9FFDAFF97FB8A20E62AB6FA49" authorityName="Dugand" authorityYear="1942" box="[1042,1172,1500,1521]" class="Magnoliopsida" family="Moraceae" genus="Ficus" kingdom="Plantae" order="Rosales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="apollinaris">
<emphasis id="B94C5E68FFDAFF97FB8A20E62AB6FA49" box="[1042,1172,1500,1521]" italics="true" pageId="2" pageNumber="3">F. apollinaris</emphasis>
</taxonomicName>
(
<emphasis id="B94C5E68FFDAFF97FB3B20E62B0EFA49" box="[1187,1324,1500,1521]" italics="true" pageId="2" pageNumber="3">
<taxonomicName id="4C38F9F9FFDAFF97FB3B20E62AD6FA49" authorityName="Beccari" authorityYear="1887" box="[1187,1268,1500,1521]" class="Liliopsida" family="Arecaceae" genus="Barbosa" kingdom="Plantae" order="Arecales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="genus">Barbosa</taxonomicName>
1245
</emphasis>
). C,D. 
<taxonomicName id="4C38F9F9FFDAFF97FAEE20E62ECAF9AD" class="Magnoliopsida" family="Moraceae" genus="Ficus" kingdom="Plantae" order="Rosales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="carchiana">
<emphasis id="B94C5E68FFDAFF97FAEE20E62ECAF9AD" italics="true" pageId="2" pageNumber="3">Ficus carchiana</emphasis>
</taxonomicName>
(
<emphasis id="B94C5E68FFDAFF97FF6E233A2F85F9AD" box="[246,423,1536,1557]" italics="true" pageId="2" pageNumber="3">McPherson 13393</emphasis>
).
</paragraph>
</caption>
<paragraph id="8B87827AFFDAFF97FF25237F2DB1F93A" blockId="2.[136,1452,1605,1810]" pageId="2" pageNumber="3">
<emphasis id="B94C5E68FFDAFF97FF25237F2F8FF9E6" bold="true" box="[189,429,1605,1630]" pageId="2" pageNumber="3">Selected specimens:</emphasis>
— 
<collectingCountry id="F32FC2EAFFDAFF97FE52237F2C40F9E6" box="[458,610,1605,1630]" name="Colombia" pageId="2" pageNumber="3">COLOMBIA</collectingCountry>
. 
<collectingRegion id="49FC4C98FFDAFF97FDE8237F2CC4F9E6" box="[624,742,1605,1630]" country="Colombia" name="Putumayo" pageId="2" pageNumber="3">Putumayo</collectingRegion>
: Mocoa, Mocoa, Camino Viejo San Antonio, 
<emphasis id="B94C5E68FFDAFF97FB67237F2B89F9E6" box="[1279,1451,1605,1630]" italics="true" pageId="2" pageNumber="3">Plowman 2030</emphasis>
(COL, 
<collectingCountry id="F32FC2EAFFDAFF97FF4F23532EDDF93A" box="[215,255,1641,1666]" name="United States of America" pageId="2" pageNumber="3">US</collectingCountry>
); San Antonio del Guamués, 
<emphasis id="B94C5E68FFDAFF97FDDE23532D3EF93A" box="[582,796,1641,1666]" italics="true" pageId="2" pageNumber="3">Cuatrecasas 11214</emphasis>
(COL, F).
</paragraph>
<paragraph id="8B87827AFFDAFF97FF2523B72D6EF8AA" blockId="2.[136,1452,1605,1810]" pageId="2" pageNumber="3">
<emphasis id="B94C5E68FFDAFF97FF2523B72F04F91E" bold="true" box="[189,294,1677,1702]" pageId="2" pageNumber="3">Notes:—</emphasis>
<taxonomicName id="4C38F9F9FFDAFF97FEBE23B72FDDF91E" authorityName="Dugand" authorityYear="1942" box="[294,511,1677,1702]" class="Magnoliopsida" family="Moraceae" genus="Ficus" kingdom="Plantae" order="Rosales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="aequatorialis">
<emphasis id="B94C5E68FFDAFF97FEBE23B72FDDF91E" box="[294,511,1677,1702]" italics="true" pageId="2" pageNumber="3">Ficus aequatorialis</emphasis>
</taxonomicName>
was synonymized under 
<taxonomicName id="4C38F9F9FFDAFF97FC8323B72AB8F91E" authority="Standley (1937: 305)" authorityName="Standley" authorityPageNumber="305" authorityYear="1937" box="[795,1178,1677,1702]" class="Magnoliopsida" family="Moraceae" genus="Ficus" kingdom="Plantae" order="Rosales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="macbridei">
<emphasis id="B94C5E68FFDAFF97FC8323B72D8AF91E" box="[795,936,1677,1702]" italics="true" pageId="2" pageNumber="3">F. macbridei</emphasis>
<bibRefCitation id="EFA9FF8BFFDAFF97FC3623B72AB8F91E" author="Standley, P. C." box="[942,1178,1677,1702]" pageId="2" pageNumber="3" pagination="384 - 388" refId="ref21019" refString="Standley, P. C. (1937) Flora of Costa Rica 18, part 1: 384 - 388." type="book chapter" year="1937">Standley (1937: 305)</bibRefCitation>
</taxonomicName>
by 
<bibRefCitation id="EFA9FF8BFFDAFF97FB5B23B72B48F91E" author="DeWolf, G. P." box="[1219,1386,1677,1702]" pageId="2" pageNumber="3" pagination="1 - 20" refId="ref19937" refString="DeWolf, G. P. (1965) Ficus, subgenus Pharmacosycea in America. Elliottia 4: 1 - 20." type="journal article" year="1965">Dewolf (1965)</bibRefCitation>
, 
<bibRefCitation id="EFA9FF8BFFDAFF97FAED23B72FB2F972" author="Berg, C. C. &amp; Villavicencio, X." pageId="2" pageNumber="3" pagination="1 - 173" refId="ref19384" refString="Berg, C. C. &amp; Villavicencio, X. (2004) Taxonomic studies on Ficus (Moraceae) in the West Indies, extra-Amazonian Brazil and Bolivia. Ilicifolia 5: 1 - 173." type="journal article" year="2004">Berg &amp; Villavicencio (2004)</bibRefCitation>
and 
<bibRefCitation id="EFA9FF8BFFDAFF97FE53238B2C7AF972" author="Berg, C. C." box="[459,600,1713,1738]" pageId="2" pageNumber="3" pagination="7 - 48" refId="ref19134" refString="Berg, C. C. (2009) Moraceae (Ficus). In: Harling, G. &amp; Persson, C. (Eds.) Flora of Ecuador, vol. 85. University of Gothenburg, Goteborg, pp. 7 - 48." type="book chapter" year="2009">Berg (2009)</bibRefCitation>
, but we observed that the former species differs from the latter by stipule persistence (subpersistent in 
<emphasis id="B94C5E68FFDAFF97FE4B23EF2C92F956" box="[467,688,1749,1774]" italics="true" pageId="2" pageNumber="3">
<taxonomicName id="4C38F9F9FFDAFF97FE4B23EF2CA5F956" authorityName="Dugand" authorityYear="1942" box="[467,647,1749,1774]" class="Magnoliopsida" family="Moraceae" genus="Ficus" kingdom="Plantae" order="Rosales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="aequatorialis">F. aequatorialis</taxonomicName>
vs.
</emphasis>
caduceus in 
<taxonomicName id="4C38F9F9FFDAFF97FCD223EF2DF8F956" authorityName="Standley" authorityYear="1937" box="[842,986,1749,1774]" class="Magnoliopsida" family="Moraceae" genus="Ficus" kingdom="Plantae" order="Rosales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="macbridei">
<emphasis id="B94C5E68FFDAFF97FCD223EF2DF8F956" box="[842,986,1749,1774]" italics="true" pageId="2" pageNumber="3">F. macbridei</emphasis>
</taxonomicName>
), lateral bracts (absent or isolated 
<emphasis id="B94C5E68FFDAFF97FAF523EF2BAEF956" box="[1389,1420,1749,1774]" italics="true" pageId="2" pageNumber="3">vs.</emphasis>
in whorls) and distribution (
<collectingRegion id="49FC4C98FFDAFF97FE3C23C32C77F8AA" box="[420,597,1785,1810]" country="Colombia" name="Cauca" pageId="2" pageNumber="3">Cauca province</collectingRegion>
<emphasis id="B94C5E68FFDAFF97FDC323C32C58F8AA" box="[603,634,1785,1810]" italics="true" pageId="2" pageNumber="3">vs.</emphasis>
Yungas province).
</paragraph>
</subSubSection>
</treatment>
</document>