The recent apple snails of Africa and Asia (Mollusca: Gastropoda: Ampullariidae: Afropomus, Forbesopomus, Lanistes, Pila, Saulea): a nomenclatural and type catalogue. The apple snails of the Americas: addenda and corrigenda
Author
Cowie, Robert H.
text
Zootaxa
2015
3940
1
1
92
journal article
10.11646/zootaxa.3940.1.1
5013eaff-bb37-4be3-a1ea-43c11d0cf5d1
1175-5326
288204
B5D45D1B-4B52-4F0B-8AF6-B587F8857475
intortus
Ampullaria intorta
Lamarck, 1816
: pl. 457, fig. 4a, b [1822a: 179].
Type
material
—
syntype
: MHNG-INVE- 51297 (formerly 1093/100).
Type
locality
—no locality given.
Distribution
—
Gambia
(
Paetel 1887: 479
),
Democratic Republic of the Congo
(
Dupuis & Putzeys 1902: LX
), Lower
Congo
River (
Pilsbry & Bequaert 1927: 192–193
).
Remarks
.
Mermod (1952: 92–95)
discussed the single specimen in the Lamarck collection in MHNG, considering it to be “le
Type
” and that it is faithfully represented by the illustrations of Lamarck (1816: pl. 457, fig. 4a, b).
Binder (1957: 108)
also referred to it as “Le
type
”. However, neither citation of “
Type
”/”
type
” is necessarily evidence that the specimen is a
holotype
(
Code
, Art. 72.4.7), they do not constitute
lectotype
designations (
Code
Arts. 74.5, 74.6) and, following the
Code
(Rec. 73F), a
holotype
should not be assumed. Therefore, it is here treated as a
syntype
, pending possible future designation as a
lectotype
.
Binder (1957: 107–109)
discussed the confused history of this species, arising primarily from the supposed misidentification of Lamarck’s species by
Martens (1879: 191, pl. 157, figs. 1–3)
, Kobelt (1912b: 27, pl. 29, figs. 4–6) and
Pilsbry & Bequaert (1927: 192, pl. 18, figs. 2–4)
, and from the identification by
Morelet (1848: 354)
of his specimen of
intorta
Lamarck
as a distinct species that he named
libyca
. To maintain stability of the usage of
intorta sensu
Martens
and of
libyca
Morelet, Binder
suppressed
intorta
Lamarck
as a junior synonym of
libyca
Morelet. This
treatement was followed by
Brown (1980
: 49, 1994: 59) and
Berthold (1991: 246)
, and by
Köhler & Glaubrecht (2006: 206)
, who designated a
lectotype
(ZMB 39407a) and
paralectotypes
(ZMB 39407b (2 spms.), ZMB
111747
(1 spm.)) for
intorta sensu
Martens. However
, according to the
Code
(Art. 49) a “previously established specific or subspecific name wrongly applied to denote a species-group taxon because of misidentification cannot be used as an available name for that taxon”. Hence,
intorta
Martens
is unavailable and another name must be used for the taxon he actually treated, if indeed it is distinct from
intorta
Lamarck. The
lectotype
and
paralectotypes
designated by
Köhler & Glaubrecht (2006: 206)
have no status. In addition, if
libyca
Morelet
is in fact not distinct from
intorta
Lamarck
, it must be treated as a junior synonym of
intorta
Lamarck.
Binder’s interpretations require verification. Therefore this
catalogue
does not assess the status of
intorta sensu
Martens
as either the same as
intorta
Lamarck
or a misidentification of his material as
intorta
Lamarck
; and it treats
libyca
Morelet
as distinct from
intorta
Lamarck
, following
Pilsbry & Bequaert (1927: 187, 192)
. Valid species,
teste
G.B. Sowerby III (1916: 66) and
Pilsbry & Bequaert (1927: 192)
, who both placed it in
Lanistes
s
.
str
.
inversa
Ampullaria inversa
Jay, 1836
: 47
. Unavailable name:
nom
.
nud
.
Remarks
. Name attributed to “Lam.” (= Lamarck) and the material stated as originating from
Egypt
. Listed here under
Lanistes
, as the name
inversa
probably reflects the superficially sinistral shell of this genus.