' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae) Author Jałoszyński, Paweł text Zootaxa 2014 3828 1 1 76 journal article 36382 10.11646/zootaxa.3828.1.1 8c1cda9b-0d82-41f3-a1c1-d8eb574a1ba7 1175-5326 286513 B2FEAE60-7E51-45FA-A38F-930A084A5AAA Scydmaenozila mallacootana sp. n. ( Figs. 74 , 85–86 , 93–94 , 195 c) Type material studied. Holotype : AUSTRALIA : ♂: four labels: " AUSTRALIA / Victoria / Mallacoota N.P. / 26 May 1978 / S. & J. Peck" [white, printed], "mushroom / litter' [white, printed], "Aust. Nat. / Ins. Coll." [green, printed], " SCYDMAENOZILA / mallacootana m. / det. P. Jałoszyński, 2014 / HOLOTYPUS " [red, printed] ( ANIC ). Diagnosis. Male and female: body uniformly brown. Male: aedeagus slender, in ventral view with broadly rounded apex; apical part of median lobe abruptly demarcated. Description. Body of male ( Fig. 74 ) strongly convex, elongate and slender, with long appendages, BL 2.75 mm ; cuticle glossy, dark brown with reddish hue, elytra slightly lighter than head and pronotum, vestiture dark brown. Head ( Fig. 74 ) broadest at eyes, HL 0.50 mm , HW 0.60 mm ; tempora longer than eyes, strongly curved posteromesally; vertex weakly convex; frons subtrapezoidal. Eyes large and strongly projecting laterally from the head silhouette, finely faceted. Punctures on head dorsum fine, separated by spaces 1.5– 2 x as wide as diameters of punctures; setae on frons and anterior portion of vertex short, sparse and suberect, tempora and posterior portion of vertex densely covered with long bristles directed posteriorly and posterolaterally. Antennae ( Fig. 74 ) slender, AnL 1.65; antennomeres I–VII distinctly elongate, VIII slightly longer than wide, IX–X each about as long as broad, XI as long as IX–X together, 2.5x as long as broad, pointed. Pronotum ( Fig. 74 ) in dorsal view strongly elongate, broadest near anterior third, PL 0.80 mm , PW 0.63 mm ; anterior and lateral margins in anterior half rounded; sides slightly concave in posterior third; posterior margin distinctly bisinuate; base of pronotum with two pairs of large and deep pits, additionally with a pair of shallow but distinct impressions located dorsolaterally. Punctures on pronotal disc fine and inconspicuous; entire dorsal and lateral surface of pronotum covered with moderately dense bristles, dorsally with admixture of thin setae. Elytra ( Fig. 74 ) slightly rhomboidal, broadest near middle, EL 1.45 mm , EW 1.05 mm , EI 1.38; basal impressions short but distinct; elytral apices separately rounded. Punctures on elytral disc fine but slightly more distinct than those on pronotum, separated by spaces 2– 3 x as wide as diameters of punctures; setae long, curved, sparse and erect. Hind wings well developed, about twice as long as elytra. Legs ( Fig. 74 ) long and slender, without modifications. Aedeagus ( Figs. 85–86 , 93–94 ) elongate and relatively slender; AeL 0.63 mm ; in ventral view short apical part abruptly demarcated, with broadly and weakly rounded apical margin; in lateral view apical part subtriangular and sharply demarcated from ventral wall; internal armature complicated, with submedian complex of elongate and darkly sclerotized structures and subapical finely denticulate lateral lobes of internal sac; parameres long and slender, with distinctly broadened apices. Female . Unknown. Distribution ( Fig. 195 c). South-eastern Australia : Victoria. Etymology. Locotypical, after the Mallacoota National Park, where this species occurs. Remarks. Scydmaenozila mallacootana has the aedeagus very similar to that of Scydmaenozila macrosticta ; differences can be seen only in the arrangement of internal structures. However, externally these species are distinctly different: Scydmaenozila macrosticta has the bicolorous pattern on elytra, while Scydmaenozila mallacootana is uniformly brown; also the shape of the pronotum and elytra is different ( Fig. 72 vs. 74).