A review of the genera Myotis, Ia, Pipistrellus, Hypsugo, and Arielulus (Chiroptera: Vespertilionidae) from Myanmar (Burma), including three species new to the country Author Bates, Paul J. J. Harrison Institute Centre for Systematics and Biodiversity Research Bowerwood House St Botolph’s Road Sevenoaks Kent TN 13 3 AQ Great Britain; E-mail: hzm @ btinternet. com hzm@btinternet.com Author Nwe, Tin Department of Zoology University of Yangon Yangon Myanmar Author Bu, Si Si Hla Department of Zoology University of Yangon Yangon Myanmar Author Mie, Khin Mie Department of Zoology University of Yangon Yangon Myanmar Author Swe, Khin Maung Department of Zoology University of Yangon Yangon Myanmar Author Nyo, Nyo Zoology Department University of Veterinary Science Yezin Myanmar Author Khaing, Aye Aye Department of Zoology University of Yangon Yangon Myanmar Author Aye, Nu Nu Department of Zoology University of Yangon Yangon Myanmar Author Toke, Yin Yin Department of Zoology Dawei University Tanintharyi Division Myanmar Author Aung, Naing Naing Department of Zoology Hinthida University Ayeyarwady Division Myanmar Author Thi, Mar Mar Zoology Department, University of Distance Education, Yangon, Myanmar Author Mackie, Iain Harrison Institute, Centre for Systematics and Biodiversity Research, Bowerwood House, St Botolph’s Road, Sevenoaks, Kent TN 13 3 AQ, Great Britain text Acta Chiropterologica 2005 2005-09-01 7 2 205 236 journal article 10.3161/1733-5329(2005)7[205:AROTGM]2.0.CO;2 1733-5329 3942897 Myotis montivagus Burmese Whiskered Bat Vespertilio montivagus Dobson, 1874: 237 ; Hotha, Yunnan , China TABLE 5. Extended

Species	C–M3	M3–M3	C–M3	C1–C1	M
Myotis mystacinus	5.4	5.3	5.6	3.4	10.1
M. muricola	4.8, 5.1	5.4, 5.5	5.1, 5.4	3.3, 3.6	9.2, 9.8
M. siligorensis	4.4 (1)	4.7 (1)	4.7 (1)	2.2 (1)	8.4, 8.6
4.5	4.8	5.5	?	8.9
M. montivagus	6.2	6.8	6.4	4.3	11.7
6.1	6.8	6.5	4.3	11.7
M. hasselti	6.0, 6.2	6.2, 6.5	6.6, 6.9	4.4, 4.6	11.9, 12.1
M. horsfieldi	5.5, 5.6	6.2, 6.3	6.1, 6.3	4.2, 4.3	10.8, 10.9
M. chinensis	9.7, 0.2	9.6, 0.3	10.3, 0.3	6.4, 0.3	18.2, 0.6
9.5–9.9	9.2–10.0	10.1–10.8	6.2–6.8	17.8–19.2
9.5	9.4	10.3	6.3	17.8
Ia io	10.9	11.9	11.8	9.0	21.5
Pipistrellus paterculus	4.2, 0.2	5.5, 0.2	4.3–4.4 (2)	3.9–4.2 (2)	9.3–9.7 (2)
4.0–4.4	5.2–5.7
P. ceylonicus	5.5	6.6	5.9	5.0	11.2
P. javanicus	4.8, 0.1	5.9, 0.1	5.1, 0.3	4.5, 0.2	9.5, 0.2
4.6–5.0 (12)	5.6–6.1 (12)	4.2–5.5 (12)	4.2–4.7 (12)	9.2–9.9 (11)
4.8	6.0	5.4	4.4	9.5
P. abramus i	4.2 , 4.3	5.2 , 5.4	4.5 , 4.6	3.8 , 4.0	8.8 , 9.0
P. coromandra	4.3	5.3	4.6	3.9	8.6
P. tenuis	3.7, 0.1	5.2, 0.1	4.0, 0.1	3.7, 0.2	7.7, 0.2
3.7–3.8	5.1–5.4	4.0–4.1	3.5–3.8	7.4–8.0
P. pulveratus	5.0, 0.1	5.8, 0.2	5.5, 0.2	4.4, 0.2	9.9, 0.3
4.7–5.3	5.6–6.2	5.1–5.8	4.1–4.7	9.5–10.4
P. lophurus	4.8	6.2	5.2	4.9	10.0
P. cadornae ii	4.6, 4.7	5.8, 5.8	4.9, 5.1	4.5, 4.6	8.5, 9.8
4.6 (1)	5.9 (1)	5.0 (1)	4.8 (1)	10.0 (1)
P. joffrei	5.1	7.0	5.4	5.2	10.8
5.0 (1)	7.1–7.2 (2)	5.5, 0.1	–	11.0, 0.2
5.4–5.6	10.8–11.2
P. anthonyi iii	5.3	7.2	–	5.1	–
P. affinis	5.3, 5.3	6.1, 6.3	5.8, 6.0	4.9, 4.9	10.5, 10.6
Arielulus circumdatus iv	6.3 (1)	7.5 (1)	7.2 (1)	5.2 (1)	12.2 (1)

i — measurements of extralimital specimens from Vietnam (HZM specimens) ii — measurements (external only) of extralimital specimens from Vietnam ( Bates et al ., 1997 ) iii — measurements based on Tate (1942) iv — measurements of extralimital specimens from India and Cambodia (BMNH and HZM specimens) Previous records from Myanmar Kachin State : Adung Valley ( Hill, 1962 ). There is no new material. Descriptive characters This is a medium-large Myotis with a forearm length 40.5, 40.6 mm for the two specimens known from Myanmar ( Table 4 ; FA = 44.7–46.8 mm in Bates and Harrison, 1997 ; 44.0– 44.6 mm in Hendrichsen et al. , 2001 , and 39–47 mm in Borissenko and Kruskop, 2003 ). The dorsal pelage is dark brown with paler tips that give a slight glossy sheen. On the ventral surface, the hair bases are dark and the tips paler brown to fawn. In the wings, the third metacarpal exceeds the fourth and fifth in length. The wings are attached to the base of the outer phalanx of each foot. The feet are not en- larged, equal to or less than half the length of the tibiae. The skull is robust. The second upper premolar (P3) appears to be variable in size. In extralimital material, it is be- tween one eighth and one third the crown area of the first (P2) and is displaced inter- nally; P2 and the third premolar (P4) are in contact in some individuals. The lower ca- nine (C1) is short, only slightly exceeding the third lower premolar (P4) in height. The second lower premolar (P3) is half the crown area of the first (P2), and is com- pressed tightly in the toothrow; occasional- ly, it is displaced internally. Similar species Myotis montivagus is comparable in forearm length to M. hasselti and M. hors- fieldi but differs in the smaller size of its feet relative to tibia length. Taxonomic notes Corbet and Hill (1992) noted that the specimens from Myanmar and southern China are relatively small when compared to referred material from India and else- where in South-East Asia. The taxonomic significance of this is not known. Ecological notes In Kachin State , it was collected in 1931 at an altitude of 1,850 m ( 6,000 feet ) in the Adung Valley, which is in the foothills of Mount Khakaborazi in the eastern Himalaya ( Hill, 1962 ).