Fifty new genera of Hesperiidae (Lepidoptera) Author Cong, Qian Author Zhang, Jing Author Shen, Jinhui Author Grishin, Nick V. text Insecta Mundi 2019 2019-10-11 731 731 1 56 journal article 10.5281/zenodo.3677235 1ee19d5d-27cf-4c93-989c-3fe3eb467203 1942-1354 3677235 BA35690A-FC73-4E5A-A805-FE9550275FEC Flattoides Grishin , new genus http://zoobank.org/ 9D7A5387-08F8-4C25-B584-B67F90EEC43A Type species. Codatractus amazonensis Bell, 1947 . Definition. A sister genus to Oileides Hübner, [1825] ( type species Oileides vulpinus Hübner, [1825] ), where C. amazonensis was placed previously ( Fig. 2 ). However large genetic distance between O. vulpinus and O. amazonensis (COI barcodes differ by 12.3%, the difference is typically smaller than 10% for species within a genus) argues for their distinction. Keys to D. 9.4 in Evans (1952) . Genitalia illustrated by Bell in his fig. 2 ( Bell 1947 ). Distinguished from its relatives by the following combination of characters: shorter than tegumen and broad uncus with small (not longer than wide) arms; prominent gnathos not shorter than uncus; valva gradually curved dorsad, ending with two pointed teeth; tuft of apricot-colored modified scales in a groove along the vein near the base of ventral hindwing in males; forewing with a macular white band from mid-costa to tornus. In DNA, a combination of the following base pairs is diagnostic: aly8661.4.1:A916G, aly10226.44.1:C1975G, aly709.1.2: T194 C, aly36556.1.1: T2416 T (not A), aly274.33.1:A188A (not G), aly82.28.5: T398 T (not G). Etymology. The name is a masculine noun in the nominative singular, for the type species that resembles butterflies from the genus Celaenorrhinus known as “Flats” in some English-speaking countries. Species included. Only the type species. Parent taxon. Subtribe Oileidina Grishin, 2019.