Fossil butterflies, calibration points and the molecular clock (Lepidoptera: Papilionoidea) Author Jong, Rienk De text Zootaxa 2017 4270 1 1 63 journal article 32975 10.5281/zenodo.583183 6c479acc-8b18-4f0b-a6e5-85bcd6d7b6b7 1175-5326 583183 2D00AFF5-4FE2-4EC1-A328-C8670CFB8D6D reynesii . Satyrites reynesii Scudder, 1872 Nymphalidae : Satyrinae . France, Bouches-du-Rhône, Aix-en-Provence; late Oligocene–early Miocene. Depository: MVMF (holotype). Published figures: Leestmans (1983: Figs 9–10) ; Scudder (1875: Pl. I Figs 2 , 5 ). The original description ( Scudder 1872 ) is repeated by Scudder (1875), who erected a new genus for the fossil: Lethites . Kozlov (1988) accepted Scudder’s genus switch. The poorly preserved specimen bears only the forewings, which yield some usable characters. Subcosta basally strongly swollen; radial formula 1, 2, 3+(4+5), cell closed. A swollen subcosta is typical, but not universal, for Satyrinae (Nymphalidae) , but this character occurs in some genera of the Biblidinae as well ( Biblis , Ergolis , Eurytela , Hypanis , Vila ). In Nymphalinae with swollen Sc, the cell is usually open or at least, the ldc is weak. In the fossil ldc seems to be fairly strong. Thus, it is plausible to consider the fossil a member of the Satyrinae . According to Scudder it is similar to the recent genera Lethe and Cercyonis , but it is not clear why it should be more similar to these genera than to any other satyrine genus. The similarity is not an indication of relationship, since Lethe and Cercyonis are only remotely related ( Wahlberg et al . 2009 ). Moreover, in Lethe there are no basally swollen veins in the forewing ( Ackery et al . 1999 ). Neorinopsis, Pseudoneorina , Neorinella and Satyrites have been classified, without comment, as fossil genera belonging to the subtribe Lethina of the tribe Satyrini by Wahlberg & Peña (2015) , but these choices are not supported by apomorphies. For further details, see the discussion under Neorinella garciae and Neorinopsis sepulta .