Revision ofEutyphlusLeConte (Coleoptera: Staphylinidae: Pselaphinae), with Description of a New Species and Phylogenetic Placement within the Tribe Trichonychini Author Owens, Brittany E. Author Carlton, Christopher E. text The Coleopterists Bulletin 2016 2016-03-31 70 1 1 29 http://dx.doi.org/10.1649/072.070.0102 journal article 10.1649/072.070.0102 5341032 Eutyphlus similis Material Examined. ( n = 737) *TENNESSEE: Sevier Co.; GSMNP, Indian Head Tr.; 35.60944°N 83.44659°W ; Sift litter. 5290′; 20 July 2003 . S. O’ Keefe/ ( LSAM )(slide mounted male)(description of male based on this specimen). *VIRGINIA: Page Co., Shennandoah Natl. Park, Little Stony Man at 38°36.24′N 78°21.99′W . 1024m . For. litter 3 May 2006 . A.Tishechkin ( LSAM )(slide mounted female)(description of female based on this specimen). * USA , AR , Washington Co. Ozark Nat. Forest Wedington 36°06.312N 94°23.390W Litter between rocks 25 Jul. 2010 . MJ Skvarla APGD 10-0726-004 ( LSAM )(16F). *Cloudland Canyon S.Pk. Dade Co., GA . 17. VI .62 Forest floor debris/ H. R . Steeves Jr. collector/ H. R . Steeves Jr. Collection/ CNHM 1963 H. R .Steeves, Jr. Pselaphidae Colln. Acc. Z-13, 288 ( FMNH )(2F; 1M). * USA : GEORGIA , Dade Co., Cloudland Canyon State Pk. 34°48.88′N 85°29.10′W 510m . 17 Sept. 2006 . Forest litter sifting. I.M.Sokolov ( LSAM ) (6F). * GA :White Co. 8 August 1984 coll. R .J. Beshear “ Eutyphlus ”/ det. DSChandler ( GMNH ). * KY . Edmonson Co. Mammoth Cave Nat. Park Ugly Creek 8.5 mi NE Rhoda Berlese sample of rotted leaves 22-NOV-1992 , E.A. Lisowski ( LSAM ) (4F). *Mammoth Cave KY . V-3-47 O.Park/ Field. Mus. Nat. Hist. Orlando Park Pselaphidae Colln. (FMNH) (2F). * USA : MD : Calv. Co. 4 mi S Prince Frederick, IV-16/ V-7-1987 , L Masner FIT, forest by crk ( DSC )(2M). *N CAROLINA: Avery Co. Blue Ridge Pkwy., Grandfather Mt. at 36°5.70′N 81°49.62′W 1580M. Litter sifting 1 May; 2006. A.K.Tishechkin ( LSAM )(19F). *Clingman’ s Peak Buncoabe Co., N.C. 3-VII-60 spruce grove debris/ H. R . Steeves Jr. Collector/ H. R . Steeves Jr. Collection/ CNHM 1963 H. R .Steeves, Jr. Pselaphidae Colln. Acc. Z-13, 288/ Field Mus. Nat. Hist. Orlando Park Pselaphidae Colln. (FMNH) (1F). *NCBuncoabeCo.; MtPisgah5600 26Ap74PDebs soiluRocks ( LSAM )(2F). *USANCCALDWELLCO Nr Cragg 36°4′31″ N 81°46′52″ W el 2930′ 14Nov2009 J&SCornell ExMoss & Rhododendron litter RoadsideDitch ( LSAM )(1F). *Grandfather Mt. N.C. 14.X.50 T .Daggy/ Field Mus. Nat. Hist. Orlando Park Pselaphidae Colln. (FMNH) (1F). * USA : NC : Haywood GSMNP BRPW nr. Cove Field Ridge Overlook: litter/ 35°25.84′N 83°21.15′W’ 1420m 21 Sep2005 ATishechkin ( LSAM )(5F).*NCAROLINA: Haywood Co. GSMNP, Caldwell Fork Tr. At UTM 30897 E; 3940883N. Moist forest Berlese. 3 August 2002 .C.Carlton&N.Lowe( LSAM )(6F;1M).* U.S. A, NC Haywood Co. GSMNP, Balsam Mt. Trail 83°10′57″W , 35°38′12″N / 20 Oct 2001 , A. Tishechkin elv. 1500m , hdwd. Berlese ( LSAM )(3F). * U.S.A. , NC , Haywood Co. GSMNP, Baxter Creek Tr. 83°06′36″W , 35°43′24″N / elv. 1350m .forest litter, Berlese 23 March 2002 , C.Carlton V .Moseley/ LSAM 0014787 ( LSAM )(1F; 2M). * USA : NC : Haywood lower Baxter Cr. Tr/308798 3957165 5 Jun 2005 CWright/ LSAM 0092285 ( LSAM )(8F). * USA : NC : Haywood Co. GSMNP Catalooche Divide Tr. Near Purchase/ 311819E 3940339N 17 Jul 2002 CCarlton ( LSAM )(2F). * USA : NC : Haywood Co. GSMNP Cataloochee Rough Ridge Tr., lower/ 306360E 3940881N moist berlese 29 Jul 2002 CCarlton ( LSAM )(2F). * U.S.A. , NC , Haywood Co. GSMNP, Chestnut Branch Trail 83°07′24″W , 35°45′34″N elv. 740m , leaf litter Berlese 1 August 2001 , A.Tishechkin ( LSAM ) (4F). * USA : NC : Haywood Co. GSMNP McKee Branch Tr./310220E 3940349N, most upland berlese 16 Jul 2002 CCarlton ( LSAM )(1F). * USA : NC : Haywood Co.; GSMNP Purchase; Knob, berlese/313124E; 3939640N 5089 ft ; 20 July 2002 ; CCarlton ( LSAM )(1F). *MECKLENBURG Co., N.C. iv-22-57 T .Daggy ( NCSUIM )(2F; 1M). * NORTH CAROLINA : Swain Co., GSMNP, admin. Road W of Fontana Dam, Lewellyn Branch UTM 0250085E 392971N 12 Apr 2004 . A.J. Mayor ( LSAM )(1F). *N CAROLINA: Swain Co. GSMNP, Appalachian Tr. ∼ 1.5mi E Newfound Gap 35°36.96′N 83°24.66′W 1730m . Forest litter 18 July 2002 . C.Carlton ( LSAM )(10F). * USA : NC : Swain Co GSMNP Appalachian Tr. 1.5 mi NE Newfound Gap/ 281639E 3944038N ridgetop berlese 18 July 2002 CCarlton ( LSAM )(3F; 1M). * USA : NC : Swain Co. GSMNP Appalachian Tr. at Sweat Heifer Cr. Tr. Ridgetop berlese/ 282160E 3944324N 18 July 2002 CCarlton ( LSAM )(2F; 1M). *N CAROLINA: Swain Co. GSMNP, E branch of Balsam Mt. Tr. Nr. Balsam Mt. Rd; 35°37.5′N 83°12.7′W . 980m Forest litter. 26 July 2004 J.Ciegler & M.Fitzgerald ( LSAM )(1F). *N CAROLINA: Swain Co. GSMNP, W branch of Beach Gap Tr. @ 35°37.3′N ; 83°12.8′W . 975m . Forest litter. 9.vi.2005 .A.Tishechkin ( LSAM )(1F). * U.S.A. , NC Swain Co. GSMNP, Clingman’ s Dome 83°29′50″W , 35°33′34″N /elv. 2024m , upper Berlese 30 June 2001 , Ccarlton, A.Tishechkin, V . Moseley ( LSAM )(3F). * USA : NC : Swain Co. GSMNP Upper Deep Creek Trail UTM 280352E 3942040 N moist upland berlese 22July2002 CEC/ LSAM 0077201 ( LSAM )(7F; 2M). * USA : NC : Swain Co. GSMNP, Deep Creek area, Loop Tr. at Indian Creek Trail, UTM 280689 E 3928371 N, moist forest berlese C.Carlton 26 July 2002 ( LSAM )(1F). * U.S.A. , NC , Swain Co. GSMNP, Flat Creek Trail 83° 10 ′21″W , 35° 31′1 ″N / elv. 1500m , leaf litter Berlese 31 July 2001 A. Tishechkin ( LSAM )(1F). *same locality/ LSAM 0002509 ( LSAM )(1F). *same locality/ LSAM 0002510 ( LSAM )(2F). * USA : NC : Swain Co. 5.8mi. S. Heintooga Overlook, Great Smoky Mt. N.P. 1.vi.1991 C.E.Carlton ( LSAM )(1F). * USA N.Car. Swain Co. Dirt Rd. from Heintooga Overlook to Cherokee/Lot#76-102 Oct. 14, 1976 Berlesate 5000′–4500′ R .Chenowith& R . T .Allen ( LSAM )(46F; 3M). *N CAROLINA: Swain Co.; GSMNP, Jenkins Ridge Tr. ∼ 1mi from Appalachian Tr. 35°33.75′N 83°43.2′W 1540m . Forest litter. 31 Jul; 2004. A.K.Ticheshkin ( LSAM )(1F). *same locality/ LSAM 0094941 ( LSAM )(1F). *same locality/ LSAM 0094945 ( LSAM )(3F). *N CAROLINA: Swain Co. GSMNP, upper Jenkins Ridge Tr. At Gunna Cr. 35°28.3′N ; 83°42.65′W 1400m . Forest litter. 29 July 2004 . A.Tishechkin ( LSAM )(1M). *N CAROLINA: Swain Co. GSMNP, Kephart Prong Tr. at 35°35′36″N 83°21′55″W 930m . Forest litter 20 July 2003 . A.Tishechkin ( LSAM )(1F). *N CAROLINA: Swain Co. GSMNP, Lakeshore Tr. at 35°28′20″N 83°27′3″W 1580m . Forest litter 18 July 2003 . A.Tishechkin ( LSAM ) (6F). *N CAROLINA: Swain Co. GSMNP, Lost Cove Tr. @ 35°29.45′N 83°48.10′W 685m . Forest litter. 8 May 2004 . W.D.Merritt ( LSAM )(13F; 3M). *N CAROLINA: Swain co. GSMNP, McGee Spring Cmpgr., 5 site. 35°38.31′N 83°14.36′W . 1535m . Forest litter. 2 Oct 2004 . W. Merritt ( LSAM ) (1F). *N CAROLINA: Swain Co. GSMNP, ∼ 250m W Mingus Mill Parkgr. 650m . 1m 2 litter 28 Apr 1996 . Coyle collr. ( LSAM )(2F). * USA : NC ,SwainCo. 2mi. Neintooga overlook.Beech-fir berlesate 1.vi.1990 C.E.Carlton ( LSAM )(4F; 1M). *N CAROLINA: Swain Co. GSMNP, Noland Divide Tr. at 35°33′57″N 83°28′36″W 1770m .Leaf/moss mat litter 19 July 2003 . A.Tishechkin ( LSAM ) (18F; 4M). *USANCTRANSYLVANIACONr Brevard PisgahNFPinkBedsPicnic Area N 35°21′11″ W 82°43. 557′ El 3208′ 5 Aug 09. J.F.& TADCornell AspirExGillFungi ( LSAM )(3F). * USA : NC , Watauga Co. Rough Ridge Trailhead 8 mi. SW Blowing Rock elv.4200′, 2.vi.1991 ( LSAM )(2F). * USA : NC : Wilkes Co. Stone Mt. State Park, nr. Hutchinson’ s Homestead 36°23.37′N 81°2.75′W . 485m . Litter sifting. 2 May 2006 A.K.Tishechkin ( LSAM )(2F; 1M). * USA : NC : Yancy Co. B. R . Pkw. 37.4 mi. SW intersection US 221 elv. 4000′ log litter 2.vi.1991 C.E. Carlton/ ( LSAM )(1F; 2M). * Clear Creek 10mi S Lancaster, Fair.Co., Ohio v010-1974/ DSChandler sifting rotten logs ( DSC )(1M). * USA , OH , Fairfield Co. Barneby Center 11 May 1989 P. Kovarik ( LSAM )(3F). * OHIO : Lawrence Co. Vinton Furnace Experim. Forest, nr. Forest Serv. Stn. Pu. Carrionbaited pitfall 19 Aug 1995 . P.W.Kovarik ( LSAM ) (1M). * USA , PA , Somerset Co. Mt. Davis, elv. 3200 ft. Hdwd forest berlesate 16 Aug. 1989 , Carlton ( LSAM )(4F). *St. Vinc. Penn./ 12/ C.N.H.M. Colln. (F.Psota Colln. Ex H.F. Wickham Colln.)/ Field Mus. Nat. Hist. Orlando Park Pselaphidae Colln / Eutyphlus similis 12 Lee ( FMNH )(1F). *SCGreenvilleCo 5miCaesar’ s HeadStPk5Aug95 J&Scornell Ex ForestLitter& rotFungi; 995VIII-5-2 ( LSAM )(4F). * USA SC PICKENS CO RockyBottomN 35°3′32″ W 82°47′52″E 12027′15Aug O9J&Scornellsifte& Berl-20kglitteronRockyBottom Creek ( LSAM )(6F). * USA SC PICKENS CO SassafrasMtN 35°3′49″N 82°46′42″W 15Aug 2009 J&SCornellExAlba trelius CristatusinConiFor ( LSAM )(2F). *#8/ SMOKIES , TENN . DYBAS 1953/ Field Mus. Nat. Hist. Orlando Park Pselaphidae Colln. (FMNH) (13F). * USA : TN , Blount Co. Great Smoky Mt. N.P. S. of entrance. Elv. 3000′ leaf/log litter 1.vi.1991 C.E.Carlton ( LSAM )(6F). * TENNESSEE : Blount Co. GSMNP, Ace Gap Tr. @ 35°38.7′N 83°48.3′W . 600m Forest litter. 2 Aug 2004 V .Bayless& S. Gil ( LSAM )(2F). * TENNESSEE : Blount Co. GSMNP, App. Tr. At Beechnut Gap. 35°34.16′N 83°41.51′W 1510m . Forest litter sifting 13 April 2006 . A.K.Tishechkin ( LSAM )(1F; 2M). * TENNESSEE : Blount Co. GSMNP, NE corner of Cades Cove at 35°36′33″N 83°47′12″W . 640m Forest litter. 18 July 2003 . A. Tishechkin ( LSAM )(2F). * TENNESSEE : Blount Co. GSMNP, Little Bottoms Tr. @ 35°37.125′N 83°55.397′W 520m . Forest; litter. 31 Jul 2004 . J. Ciegler ( LSAM )(14F). * TENNESSEE : Blount Co. GSMNP, lower Gregory Ridge Tr. @ 35°33.5′N 83°50.5′W . 630m . For. Litter 28 July 2004 . A.K. Tishechkin ( LSAM )(43F). * TENNESSEE : Blount Co. GSMNP, Upper Long Hungry Ridge Tr. At 35°30.89′N ; 83°51.00′W 1390m . For. Litter 12 April 2006 . A.K. Tishechkin ( LSAM )(1F). * TENNESSEE : Blount Co. GSMNP, Mt. Thunderhead nr. Summit @ 35°34.1′N 83°42.5′W . 1650m . Litter 30 July 2004 . A.K. Tishechkin ( LSAM )(2F; 1M). * TENNESSEE : Blount Co. GSMNP, West Prong Campgr. 35°37.7′N 83° 42.3°W . 520m . Forest litter. 30 Jul 2004 . J Ciegler ( LSAM )(32F). *Carter Co. Tenn. #228 Copeland 12. VII .53/ Field Mus. Nat. Hist. Orlando Park Pselaphidae Colln. (FMNH) (1F). * TENNESSEE : Cocke Co. GSMNP, Albright Grove @ 35°44.11′N 83°16.78′W 970m Forest litter. 1 Aug; 2004. J. Ciegler, A. Tishechkin ( LSAM )(6F). * TENNESSEE : Cocke Co. GSMNP, Albright Grove Loop Tr. Ca. 950m . Berlese near rotten longs 19 Jul 2003 . S.O’ Keefe ( LSAM )(10F). * U.S.A. , TN , Cocke Co. GSMNP, Albright Grove Trail 83°16′45″W , 35°44′10″N /elv. 1000m , old growth Berlese 29 June 2001 , C.Carlton, A. Tishechkin, V . Moseley ( LSAM ) (10F; 1M). * TN : Cocke Co GSMNP Albright Grove Trail loop/ 35°39′42″N 83°27′46″W 1500m Carlton Mayor 6 Dec 2003 ( LSAM )(16F). * USA Tenn. Cocke Co. GSMNP Cosby Creek Trail/ Lot #76-106 Oct. 15, 1976 Berlesate R .Chenowith& R . T .Allen ( LSAM )(19F; 1M). * USA : TN :Cocke Co. GSMNP Cosby House 35°46.67N 83°12.83W 7Oct2005 M.Gimmel sifting litter ( LSAM )(5F). * U.S.A. , TN , Cocke Co. GSMNP, Cosby Ranger Station 83°12′25″W , 35°46′12″N elv. 622m , leaf litter Berlese 31 July 2001 , A. Tishechkin ( LSAM ) (2F). * USA : TN : Cocke Co. GSMNP Gabes Mnt. Tr @ Hen Wallow Fall, moist forest berlese/ 297649E 3959487N 19 Jul 2002 CCarlton, NLowe ( LSAM )(2F). * U.S.A. , TN , Cocke Co. GSMNP, Hen Wallow Falls 83°14′27″W , 35°46′22″N elv. 580m , leaf litter Berlese 31 July 2001 , A. Tishechkin ( LSAM )(5F). * USA TN CockeCo GSMNP MaddronBaldTrail17-2916 31E3959236NA027 2320 8Aug06JCornell&Sranger BerlExSiftLitter Of30″DiamHem RockStumpJC006-VII-0-40 ( LSAM )(3F). * TENNESSEE : Cocke Co. GSMNP, Maddron Bald Tr. @ lower end of the Albright Grove Loop. ∼ 970m . Forest litter. 19 Feb 2004 . A.Mayor ( LSAM )(7F). * USA : TN : Cocke Co. GSMNP Snake Den Ridge Trail/ btw 299077 3957780 & 299657 3957885 Cwright June 2005 ( LSAM )(1F). * TENNESSEE : Sevier Co. GSMNP, App. Tr. ∼ 1mi E Newfound Gap. 35°36.96′N ; 83°24.88′W . 1650m . Forest; litter. 7 Oct 2005 D. Chouljenko& M. Gimmel ( LSAM )(1 F). * TENNESSEE : Sevier Co. GSMNP, App. Tr. At Sweat Heifer Cr. Tr. 35°37.29′N 83°24.25′W . 1725m . Forest litter. 23 Aug 2006 . V .Bayless ( LSAM )(8F; 1M). * TENNESSEE : Sevier Co. GSMNP, App. Tr. Just W of Sweat Heifer Cr. Trhd. 1720m 35°37.3′N 83°24.35′W . Forest litter. 14 April 2006 . C.Carlton ( LSAM )(3F; 1M). * U.S.A. , TN Sevier Co. GSMNP, Chimneys Picnic Area Nature Trail, 83°29′45″W , 35°38′6″N , elv. 891m /soil litter berlese 30 June 2001 , C.Carlton A.Tishechkin, V .Moseley ( LSAM )(2F). *Sevier Co. Tenn. Smoky Mt. Pk. Summer 1953 T .P. Copeland/ Field Mus. Nat. Hist. Orlando Park Pselaphidae Colln. (FMNH) (1F). *Cove Forests S.&E. Gatlinburg, TENN . Sevier Co. V-16-23 1972 2500-3500′/ Eutyphlus similis LeConte det. DSChandler ’95 ( FMNH )(1F). *Rocky Creek Gap Smoky Mts. Nat. Pk. Tenn. IX:14:53, SA-HD/ Field Mus. Nat. Hist. Orlando Park Pselaphidae Colln. (FMNH) (3F). *Gt. Smoky Mts. Nat. Pk. Elkmont Sevier Co., Tenn. 8-VI-60 Allt. 2250′ Leaf Duff/ H. Buter J. Wagner Collectors/ H. R . Steeves, Jr. Collection/ CNHM 1963 H. R . Steeves, Jr. Pselaphidae Colln. Acc. Z-13, 288/ Field Mus. Nat. Hist. Orlando Park Pselaphidae Colln. (FMNH) (2F)(2M). *Greenbrier Cove Smoky Mts. Nat. Pk. Sevier Co., TENN ./ Field Mus. Nat. Hist. Orlando Park Pselaphidae Colln. (FMNH) (8F). * USA : TN : Sevier Co. GSMNP Greenbrier Ridge Tr. 26014. 3941340. 1260m Berlese 7 Jun 2005 A. Tishechkin ( LSAM )(2F). * TENNESSEE : Sevier Co. GSMNP, Indian Head Tr. 35.60944°N 83.44659°W sift litter. 5290′ 20 July 2003 . S.O’ Keefe ( LSAM )(18F; 9M). * U.S.A. , TN , Sevier Co. GSMNP, Laurel Falls Trail 83°35′36″W , 35°40′19″N / elv. 747m , leaf litter berlese 1 July 2001 , C.Carlton, V . Moseley, A.Ticheshkin ( LSAM )(3F). * TENNESSEE : Sevier Co., GSMNP Middle Prong Tr. at 35°36.9′N 83°39.4′W 30 July 2004 . J.Ciegler ( LSAM ) (3F). * TENNESSEE : Sevier Co. GSMNP, lower Middle Prong Tr. @ 35°36.9′N 83°39.4′W . 700m . Forest litter. 30 Jul 2004 . J.Ciegler ( LSAM )(8F). * TENNESSEE : Sevier Co. GSMNP, Mt LeConte nr. Cliff Top Cabins 35°39′20″N 83°26′40″W . 1980m . Leaf moss litter. 18. VII .2003 A.Mayor& P.Skelly ( LSAM )(3F; 2M). *Newfound Gap 16.IX. 53 HSD/ Field Mus. Nat Hist. Orlando Park Pselaphidae Colln (FMNH) (2F)(1M). * USA Tenn.- N.Car.; Sevier Co.; GSMNP Newfound; Gap to Clingmans Dome/Lot #76-107; Oct. 11, 1976 ; Berlesate; R . Chenowith& R . T .Allen ( LSAM )(5F). * U.S.A. , TN , Sevier Co. GSMNP, 0.5km NE Newfound Gap, elv. 1600m 83°24′46″W , 35°38′9″N / forest litter Berlese 26 June 2001 C.Carlton, V .Moseley, A.Tishechkin ( LSAM )(1F).* TENNESSEE : Sevier Co. GSMNP, W Old Settlers Trhd. 35°42.46′N 83°22.9′W Forest litter. 6 Oct 2005 D. Chouljenko, M. Gimmel & A.Hepperman ( LSAM )(2F). * USA :, TN , Sevier Co. GSMNP, Porters Crk. Tr. 83°23′52″W , 35°40′13″N / elv. 870m , 18Oct. 2001 old growth berlese A.Tishechkin, A.Cline ( LSAM )(6F). * TENNESSEE : Sevier Co;. GSMNP, Porters Creek Tr. @ 35°40.1′N 83°23.6′W 850m . Forest litter. 31 July 2004 . C.E.Carlton&N.Lowe ( LSAM )(3F). * TENNESSEE : Sevier Co. GSMNP, Porters Creek Tr. Sift litter near rotten logs Beetle Blitz-2003 18 July 2003 . S.O’ Keefe ( LSAM )(44F). * USA : TN : Sevier Co. GSMNP Rainbow Falls snowy litter berlese/ 35°39′42″N / 83°27′46″W 1500m Carlton Mayer 6 Dec 2003 ( LSAM )(1F). * TENNESSEE : Sevier Co. GSMNP Ramsey Cascades Tr.@ 38°42.53′N 83°19.21′W 1165m . Forest litter. 10 May 2002 . W.D.Merritt ( LSAM )(3F). * U.S. A, TN , Sevier Co., GSMNP Ramsey Cascade Trail(middle) 83°18′24″W , 35°42′34″N /elv. 1300m . Forest litter berlese 21 March 2002 CECarlton, VLMoseley ( LSAM )(2F). * U.S. A, TN , Sevier Co., GSMNP Ramsey Cascade Trail(upper) 83°18′ 7″W , 35°42′30″N /elv. 1375m . Forest litter berlese 21 March 2002 CECarlton, VLMoseley ( LSAM ) (3F). * TENNESSEE : Sevier Co. GSMNP, Road Prong Tr. at 35°36′36″N 83°27′3″W 1580m . Leaf/ moss mat litter. 20 July 2003 A.Ticheshkin ( LSAM ) (20F; 1M). * USA : TN :Sevier Co. GSMNP, Roaring Fork Area, Rainbow Falls Tr./ 1Aug2002 moist cave Berlese UTM275790E 3949640N C.Carlton ( LSAM )(21F). * TENNESSEE : Sevier Co. GSMNP Trillium Gap Tr. @ 35°39.9′N 83°26.2′W 1400m . Forest litter. 29 July 2004 . A. Tishechkin ( LSAM ) (38F; 7M). TENNESSEE : Sevier Co. GSMNP, jct. Trillium Gap & Brushy Mt. Tris. 35°40.4′N 83°26.0′W . 1430m . Forest litter. 29 July 2004 C.E. Carlton & J.Ciegler ( LSAM )(16F). * VA ., Giles Co. Jefferson Nat’ l. Forest Mt. Lake Tech. Area 4. VII .67/ Rhododendron duff/ H. R .Steeves, Jr. H. R . Steeves III Collectors/ H. R .Steeves, Jr. Collection ( FMNH )(1F; 1M). * VIRGINIA : Madison Co., Shenandoah NP, nr. Upper Hawksbill Parking. 38°32.73′N ; 78°23.64′W . 1115m . Litter sifting. 3.v.2006 . A. Tishechkin ( LSAM )(2F). * Mountain Lake, VA ./ VII-23-47 J.H.Davis/ Field Mus. Nat. Hist. Orlando Park Pselaphidae Colln. (FMNH) (1F). * VIRGINIA : Smyth Co., Lewis Fork Wildrns., Mt. Rogers Tr. at 36°39.63′N 81°33.11′W 1540m . Litter sifting. 30 April 2006 . A.K.Tishechkin ( LSAM )(13F; 1M). * USA , VA . Smythe Co. 7.5; mi SE Marion, Raccoon; Branch cmpgrnd, v-29-91; sift pine & oak litter; S. O’ Keefe collection ( DSC )(1M). * USA : VA , Wash. Co. E of Damascus leaf litter 3.vi.1991 C.E.Carlton ( LSAM )(3F). * USA WV Fayette Co. 35mi N Beckley18-III-83 JFcornell ExConiferlitter JFC983-III-18-4 ( LSAM )(1M). * W. Virginia : Mercer Co. Camp Ck. St. Forest. IV:8:1971. Leg: S. Bird. FM ( HD )#71-72/ Eutyphlus similis det. DSChandler ( FMNH )(1F; 1M). * WVPocahontasCo Snedegar’ sCave (illegible) 10-vi-1989 JPCornall (illegible) forest litter ( NCSUIM )(1F; 6M).