A Checklist and New Species ofEleodesEschscholtz (Coleoptera: Tenebrionidae) Pertaining to the SubgenusPromusLeconte, with a Key to United States Species
Author
Johnston, M. Andrew
text
The Coleopterists Bulletin
2015
2015-03-31
69
1
11
19
http://dx.doi.org/10.1649/0010-065x-69.1.11
journal article
10.1649/0010-065X-69.1.11
5331184
52ED5896-72C3-4D1F-B12D-E7A02FDB7346
Eleodes
(
Promus
)
compositus
Casey 1891
, revised status
(
Fig. 1
)
The well-known coleopterist Thomas L. Casey described 15 species of
Eleodes
, of which four are currently considered valid.
Eleodes compositus
was described in 1891 and subsequently synonymized by
Blaisdell (1909)
with
Eleodes hispilabris
(Say)
, albeit without reexamining the
type
or any specimens that could be ascribed to this newly designated infrasubspecific
forma composita
(
Blaisdell 1909
). Blaisdell assessed that the aberrant form of the pronotum which Casey described “is no proof that it is specifically distinct. Analogous aberrations are observed in
dentipes
in particular, and
hispilabris
is fully as variable as that species” (
Blaisdell 1909
). Indeed, the pronotum of
E. compositus
(
Fig. 1
) is subquadrate, evenly arcuate laterally, and only slightly constricted posteriorly unlike the pronotum of
E. hispilabris
, which is distinctly constricted posteriorly, widest anterior of midline, and with very prominent anterior projections. The name
E. compositus
has since been listed as a synonym of
E. hispilabris
in several publications and its status has not been reconsidered in the literature (
Papp 1961
;
Tanner 1961
).
Fig. 1.
Eleodes compositus
. A) Holotype male, lateral habitus, scale bar = 1 cm, B) Dorsal habitus.
As part of an ongoing effort to reassess every type specimen within the genus
Eleodes
, the type of
Eleodes compositus
, residing at the USNM, was examined. In doing so, it became apparent that this specimen did not represent
E. hispilabris
, nor was it even a member of the subgenus
Eleodes
where
E. hispilabris
is placed. In fact, this specimen belongs to the subgenus
Promus
as clearly evidenced by the
holotype
male having the combined characters of spined profemora and the first two segments of the protarsi with pads of tomentose setae, a combination found in no other subgenus. The heavily sulcate elytral striae clearly distinguish this species from all other described
Promus
species.
The
holotype
, with the locality given as “
Texas
”, remains as the only known specimen of this enigmatic species.
Eleodes compositus
Casey
is hereby returned to valid, species-level standing and placed in the subgenus
Promus
.