New brachypterous species of Dichotomius Hope, with taxonomic notes in the subgenus Luederwaldtinia Martínez (Coleoptera: Scarabaeidae: Scarabaeinae) Author Nunes, Rafael V. Author Vaz-De-Mello, Fernando Z. text Zootaxa 2013 2013-01-31 3609 4 411 420 journal article 10.11646/zootaxa.3609.4.3 efd85aa0-d2a5-4d72-8125-6d45ab6935fc 1175-5326 224051 4D73C473-2533-4E1B-A039-113DF838F42A Dichotomius sexdentatus (Luederwaldt) and D. simplicicornis (Luederwaldt) Luederwaldt (1936) created the D . calcaratus species group to accommodate D. calcaratus (Arrow) , D. sexdentatus (Luederwaldt) , and D. simplicicornis (Luederwaldt) . Vulcano et al . (1976) transfered D. calcaratus to the D . buqueti species group (then in the subgenus Dichotomius ). With the transfer of the nominate species to another species group, the two remaining species, D. sexdentatus and D. simplicicornis should be placed in different species groups. After examining the holotype (MZSP), we place D. sexdentatus in the D . crinicollis species group for the following reasons: it has a tooth on clypeal genal junction ( Fig. 2 B) and its hypomeron bears dense, long and curved, red pilosity ( Fig. 2 D). After examining the holotype of D. simplicicornis (located in the NHML) and scrutinizing the original description, we place this species in the D . agenor species group based on the cephalic horn in males constituting a transverse carina with a strong central tooth and two lateral teeth ( Fig. 2 H), which is a shared characteristic with other species in this species group.