Revalidation of the Brazilian Atlantic Forest dung beetle species Coprophanaeus (Metallophanaeus) machadoi (Pereira & d’Andretta, 1955) (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini) based on morphological and distributional evidence Author Cupello, Mario Author Vaz-De-Mello, Fernando Z. text Zootaxa 2014 3869 4 435 451 journal article 10.11646/zootaxa.3869.4.7 aaa0b363-57d7-4b61-9167-7c1462dced8c 1175-5326 224348 7446EA9F-1C54-47AB-ABF1-0A82B1E59687 Placement of Coprophanaeus (Metallophanaeus) machadoi ( Pereira & d’Andretta, 1955 ) in Edmonds & Zidek’s (2010) key to species of the saphirinus group: 1. Posteromedian angle of pronotum not produced, basal fossae small, incorporated into mesial ends of basal groove. Pronotum distinctly punctured posteromedially. Paraocular areas (genae) of both sexes completely punctate, usually densely and coarsely so. Male pronotum lacking round concavities. Elytral striae carinulate, distinctly punctured ( x5 )....................................................................... Coprophanaeus (Metallophanaeus) punctatus ( Olsoufieff, 1924 ) . - Posteromedian angle of pronotum slightly produced posteriorly, very smooth, effacing basal fossae and mesial ends of basal pronotal groove. Pronotum smooth posteromedially. Male paraocular areas (genae) almost completely smooth; female paraocular area broadly rugose. Male pronotum with round concavities beneath posteromedian projections. Elytral striae simple (very rarely carinulate), very fine............................................................................. 2 2. Anterior margin of pronotum of large males with long midlongitudinal filiform prolongation, sometimes raised in sharp and acute ridge; prolongation followed by midlongitudinal well-marked carina ( Figs. 6–9 ) (in minor specimens, prolongation triangular, short or absent, carina sometimes absent). Parameres, in lateral view, weakly concave; posterior flap of apical process only slightly elevated ( Fig. 12 ). Dorsal coloration blue and green ( Figs. 1–5 ). Brazilian Atlantic Forest in eastern Minas Gerais, Espírito Santo and Rio de Janeiro (except Serra da Mantiqueira Mountains)............................................................................. Coprophanaeus (Metallophanaeus) machadoi ( Pereira & d’Andretta, 1955 ) . - Anterior margin of pronotum of large males with short triangular prolongation or rounded; pronotum smooth, midlongitudinal carina absent ( Figs. 10–11 ). Parameres, in lateral view, strongly concave; posterior flap of apical process strongly elevated and curved ( Fig. 13 ). Dorsal coloration blue, green, and red ( Figs. 15–23 ). Brazilian Atlantic Forest in Serra da Mantiqueira Mountains, southern Brazil (including São Paulo), Paraguay , and Argentina .............................................................................................. Coprophanaeus (Metallophanaeus) saphirinus ( Sturm, 1826 ) .