Type material of land snails (Mollusca: Gastropoda) described from New Zealand by taxonomists in Europe and North America between 1830 and 1934, and the history of research on the New Zealand land snail fauna from 1824 to 1917
Author
Brook, Fred J.
Author
Ablett, Jonathan D.
text
Zootaxa
2019
2019-11-14
4697
1
1
117
journal article
24883
10.11646/zootaxa.4697.1.1
2a01bfb5-6e33-42b5-ab5d-4f8512c9128f
1175-5326
3542832
AF79BEA3-3CC8-49CA-9707-A8D5B4DAACD
Helix eta
Pfeiffer, 1853
Pl. 5, fig. A
Pfeiffer, 1853. Monographia heliceorum viventium, 3: 107.
Type material:
Lectotype
,
NHMUK 1962725
(dry shell material); ex Museum Cuming (Acc. no. 1829)
.
Lectotype
fixed by inference of
holotype
(
ICZN
Article 74.6) by
Climo (1970a: 315)
(dry shell material). Possible
paralectotypes
(2), ZMH 45987, from
Altonaer Museum
, coll.
O. Semper
ex
Cuming. We
were unable to obtain images of the specimens in ZMH and do not know if they are conspecific with the
lectotype
.
Label details of
lectotype
:
‘n.sp.
H. eta Pfr. N.
Zealand’—in Pfeiffer’s handwriting.
Type
locality:
‘Nova Seelandia’ (Pfeiffer 1853: 107, 1854a: 57).
Previous illustrations of
type
material:
Climo (1981
: figs. 1A–C). A shell purported to be
Helix corniculum
[=
H. eta
, see below] was illustrated by Reeve (1852 [in
1851–1854
]: pl. 133, fig. 826) [and reproduced by
Tryon
(1887: pl. 3, fig. 12)], but does not match Reeve’s accompanying description, or the
type
material of
corniculum
, and evidently represents another species.
Remarks:
Helix eta
Pfeiffer, 1853
and
H
.
corniculum
Reeve, 1852
are based on the same
type
material, which H. Cuming had obtained from the Australian-based naturalist Frederick Strange. Pfeiffer evidently intended to publish the original description of this species in the Proceedings of the Zoological Society of London issue for 1851 (see Reeve 1852 [in
1851–1854
]: species 826;
Pfeiffer 1853a: 107
), but publication was delayed until 1854 (
Pfeiffer 1854a: 57
), and was pre-empted by Reeve (1852 [in
1851–1854
]) [note that the publication date of ‘
October 1851
’ listed for plate 133 of
Reeve (
1851
–1854) is presumed to be an error, as the immediately preceding and following plates were dated ‘
October 1852
’ (see
Petit 2007
: table 3)]. However, as noted under
corniculum
Reeve
, this name is a junior homonym of
Helix corniculum
Hombron & Jacquinot, 1847
, applied to material from New
Guinea
(
Martens 1873: 10
), and so
Helix eta
Pfeiffer, 1853
is the earliest available name for the
New Zealand
species. Historically there was considerable confusion over the identity of
Helix corniculum
Reeve
and
Helix eta
Pfeiffer
, as noted by
Climo (1981)
and
Goulstone (1995)
. Some authors (including
Suter 1913b: 725
;
Iredale 1915a: 482
;
Solem 1959: 83
;
Climo 1970a: 314
;
Powell 1979: 310
;
Schileyko 2001
: fig. 1291) used Reeve’s or Pfeiffer’s names for species with a spirally-ribbed protoconch and radially ribbed teleoconch. However,
Climo (1981)
noted that the
type
material of
Helix corniculum
Reeve
and
Helix eta
Pfeiffer
has a radially-ribbed protoconch, and matches shells that some authors (including
Suter 1913b: 715
;
Iredale 1915a: 482
;
Powell 1939: 238
;
Climo 1970a: 330
;
Powell 1979: 309
;
Schileyko 2001
: fig. 1288) had incorrectly identified as
Helix caput-spinulae
Reeve, 1852
(see entry for
caput-spinulae
below). Iredale (1915) erected the genus
Mocella
for a group of
New Zealand
charopids with spirally-ribbed protoconchs, and designated
Helix corniculum
Reeve, 1852
as the
type
species, based on a misinterpretation of that species.
Climo (1981)
and most subsequent authors have assigned
Helix eta
Pfeiffer, 1853
, and morphologically similar taxa with radially-ribbed protoconchs, to
Mocella
, based on a literal interpretation, rather than the intent, of Iredale’s (1915) generic classification (see ICZN Article 70.3.1).
Goulstone (1995: 66)
erected the genus
Climocella
for the group of
New Zealand
species with spirally-ribbed protoconchs that had previously been included in
Mocella
. New Hebridean and Solomon Island species that were assigned to
Mocella
by
Solem (1959
,
1960
) were transferred to
Sinployea
Solem, 1983
by
Solem (1983)
.
Current Taxonomy:
Listed as
Mocella eta
(Pfeiffer, 1853)
by
Climo (1981: 9)
,
Spencer & Willan (1995: 40)
and
Spencer
et al
. (2009: 215)
.
Distribution:
New Zealand
; North Island, northern South Island,
Chatham Islands
(Suter 1913b—as
Endodonta caput-spinulae
;
Wallace 1977
—as
Charopa caputspinulae
;
Powell 1979
—also as
C. caputspinulae
).