New genera, new species, and new combinations in New World Cochylina (Lepidoptera: Tortricidae: Tortricinae) Author Brown, John W. text Zootaxa 2019 2019-09-17 4671 2 195 222 journal article 25519 10.11646/zootaxa.4671.2.2 978ec4b3-8d0e-4f47-8084-610b34b96759 1175-5326 3442344 8DA2FA3F-3629-4D10-92B0-671637D91DD4 Aethes ringsi ( Metzler, 2000 ) , new combination Figs. 14 , 30 , 44 Cochylis ringsi Metzler, 2000: 191 ; Brown 2005: 213 ; Metzler & Brown 2014: 276 . Described in Cochylis by Metzler (2000) , ringsi is transferred to Aethes based on facies, DNA barcodes (see Brown et al . 2019 ), and the similarity in the male genitalia to those of Aethes tuxtlana Razowski, 1986 . The forewing pattern of A. ringsi ( Fig. 14 ) is somewhat reminiscent of that of a few other species of Aethes (e.g., A. fernaldana (Walsingham, 1879) , A. argentilimitana (Robinson, 1869)) , with a slightly shiny, buff to whitish ground color and a contrasting and characterisitic orange-brown basal patch, and a well-defined, median fascia, angled near the middle. DNA barcodes convincingly place it deeply embedded within Aethes in a maximum likelihood analyses ( Brown et al . 2019 ). The assignment of ringsi to Aethes is contradicted by the absence of the sickle-shaped processes from the socii that define Aethes and the absence of a median process of the transtilla, common to most Cochylini , including Aethes . However, at least two other species of Aethes (i.e., an undescribed species from the Midwestern U.S. and Aethes tuxtlana from Mexico ) lack the sickle-shaped processes, and the latter species lacks a median process from the transtilla, as well (see Razowski 1986b : fig. 5). The unusual somewhat ”split” valva, with a invagintation between the sacculus and the remainder of the valva of A. ringsi ( Fig. 32 ) is extremely similar to that of A. tuxtlana , which also has unusual socii. Hence, ringsi appears to be a derived species within Aethes , possibly the sister to A. tuxtlana . The female genitalia ( Fig. 44 ) are fairly typical of the genus. Aethes ringsi ranges from the southeastern U.S. ( Alabama , South Carolina , North Carolina ) to the mid-West ( Iowa , Indiana , Missouri , Arkansas , and Oklahoma ). Metzler (2000) suggests it is a species of oak barrens on sandy soils. Captures range primarily from June through August, but there are a few April and May records from the southernmost portion of its range. Holotype ³, USA , Indiana , Newton Co. , 41˚6.43’N, 87˚26.25’W, Conrad Savanna , 8 Jul 1998 , E. Metzler , UV ( USNM ).