3541 Author Judson, Sarah W. Author Nelson, C. Riley text Zootaxa 2012 2012-11-06 3541 1 118 journal article 1175­5334 505937B0-9F57-4068-82E6-8553826DD5AA Skwala pusilla (Klapalek, 1912) SYNONYMY Arcynopteryx compacta pusilla Klapálek, 1912 Arcynopteryx brevis Koponen, 1949 Arcynopterygoides vernalis Raušer, 1968 Skwala brevis Zwick, Levanidova, and Zhiltzova, 1971 Skwala pusilla Zhiltzova, 1982 TYPE LOCALITY: USSR: Krasnojarsk . DIAGNOSIS: Adults exhibit a pale stripe originating in the interocellar region and terminating on the mesonotum. The male hemitergal processes are short and blunt ( Fig. 292 ), similar to those of S. asiatica ( Fig. 288 ) and Arcynopteryx sajanensis ( Fig. 258 ). Skwala pusilla is distinguished from S. asiatica primarily by internal epiproct characters. The female subgential plate has a broad, truncate posterior margin that is indented with a slight broad depression ( Fig. 293 ), and is indistinguishable from S. asiatica ( Fig. 289 ). The nymph has a light M-line highlighted by a darker pigmentation posteriorly. The head coloration is very similar to various Arcynopteryx species ( Fig. 250 , 255 ). The pronotum is oval with a wide, dark, longitudinal stripe medially and a dark margin. DISTRIBUTION—Global: East Palearctic— Regional: AOB*, IDB— Aimag: BO, SE*, UB*. DISCUSSION: The specimens of S. pusilla documented by our survey were found near lake outlets in abundant numbers. We did not document S. pusilla in the majority of its historic range (Raušer 1968, Joost 1970, Zhiltzova 1982), in which it is sympatric with the type locality of S. asiatica .