Presence of the Glossosomatidae in Madagascar, with description of two species (Trichoptera)
Author
Gibon, François-Marie
text
Zootaxa
2017
2017-01-01
4317
1
174
178
journal article
32229
10.11646/zootaxa.4317.1.9
e3add788-8653-4f45-8658-f017da50b0b2
1175-5326
880312
Cb05909F-4E7C-4845-8C13-940B7103Aaa7
Agapetus
(
Synagapetus
)
alarum
n. sp.
Figs 1
A–1E
Holotype
:
Male
, mounted on six slides.
MADAGASCAR
:
Fianarantsoa
Prov.,
Andringitra National Park
,
Sahanivoraky River
,
22°13'33"S
,
47°0'41"E
,
810 m
a.s.l.
,
19.xi.1993
, F.-
M. Gibon
leg
. (
CBGP
).
Paratypes
:
all specimens same data as holotype;
2 males
in alcohol
; 1 male mounted on six slides; 1 male with genitalia on two slides and remaining parts in alcohol.
Diagnosis.
Agapetus alarum
n. sp.
is similar to the other Malagasy species,
A. ranohelae
n. sp.
They differ from one another by the inferior appendages (curved dorsad in
A
.
alarum
,
but rectangular in
A. ranohelae
)
and by tergum X (longer than the inferior appendages and extended in a distal digitiform lobe in
A
.
alarum
,
but as long as the inferior appendages and roughly triangular in
A. ranohelae
)
. Both species belong to the
A
. (
Synagapetus
)
agilis
Group of
Ross (1956)
. This group is not homogeneous; it includes the two southern African species:
A. agilis
, characterized by the absence of preanal appendages and the absence of a ventral process on sternite VI, and
A. murinus
, which is fairly similar to the Malagasy species but is distinguished by the slender and acute tergum X (stout and wide in the Malagasy species). Also the preanal appendages are finger-shaped in
A. murinus
, but short and wide in the Malagasy species. The other African species group (
Agapetus ungulatus
Group) is clearly characterized and distinguished by two apomorphies: a single and apically curved preapical spur on each metatibia and a highly modified tergite X.
Description.
Three ocelli. Tibial spurs 2/4/4. Maxillary palps each 5-segmented, segments 1, 2 and 5 short, segment 2 enlarged distally, segment 3 clearly longer. Labial palps each 3-segmented. Forewings each with length 4.1 mm, maximum width 1.4 mm; distal margin oval; forks 1, 2, 3, 4, and 5 present; forks 3 and 4 petiolate; Cu1 forked just beyond
m-cu
crossvein. Hindwings each with length 3.2 mm, maximum width 0.9 mm; smaller and narrower than forewing, venation reduced; forks 1, 2, 3, and 5 present; forks 1, 2, and 3 petiolate; R1 ending at R2+3; discoidal cell open; Cu1 forked just beyond
m-cu
crossvein. Sternite V with lateral fold and internal gland, similar to those described by
Ross (1956)
for
A. cocandicus
McLachlan 1875
. Sternite VI with ventral keel-shaped projection. Abdominal segment IX large and synsclerotized; tergum IX shorter than sternum. Preanal appendages short and wide, roughly rectangular. Inferior appendages each single-segmented, stout, thick, curved dorsad, with small setae but without sclerotized denticles, spines, or small bumps on inner face; 2/3 as long as tergum X. Tergum X almost entirely divided in pair of lateral elements; in lateral view each element composed of large, wide, almost rectangular basal part, ventrodistally elongated in short, acute ventral lobe and long distal finger-shaped lobe. Phallic apparatus long, slender, slightly curved ventrad.
Etymology.
From the Malagasy:
ala
(forest), with reference to the magnificent pristine forests of the Andringitra.
Habitat and distribution.
Collected from a small brook in pristine rainforest. Endemic to
Madagascar
and recorded from only Andringitra National Park (
Fig. 3
).