<document id="DE85E6BFA1646EB8B090CB80C35E8386" ID-CLB-Dataset="8656" ID-DOI="10.11646/zootaxa.4913.1.1" ID-GBIF-Dataset="bffe4ff6-abdf-4d7d-a00a-d82e333454b4" ID-ISSN="1175-5326" ID-Zenodo-Dep="4448915" ID-ZooBank="E3540F07-4CF6-4EF2-A533-64A2D49D68FF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1611043839000" checkinUser="plazi" docAuthor="Stekolnikov, Alexandr A." docDate="2021" docId="E016879A6656290233F3AA63FE621AC3" docLanguage="en" docName="zootaxa.4913.1.1.pdf" docOrigin="Zootaxa 4913 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Helenicula signata" docType="treatment" docVersion="27" lastPageNumber="87" masterDocId="1C2FFFE2660029543364AB29FF97180E" masterDocTitle="A checklist of chigger mites (Acariformes: Trombiculidae) of Southeast Asia" masterLastPageNumber="163" masterPageNumber="1" pageNumber="87" updateTime="1698924849085" updateUser="ExternalLinkService">
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<mods:title id="19A32DBC7A2CB92BFF9852D8F7E176B4">A checklist of chigger mites (Acariformes: Trombiculidae) of Southeast Asia</mods:title>
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<mods:namePart id="4A2836F8ED57409A36E7C1D840EDBC87">Stekolnikov, Alexandr A.</mods:namePart>
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<heading id="334881E06656290233F3AA63FDC1196A" bold="true" box="[151,598,330,357]" fontSize="11" level="2" pageId="86" pageNumber="87" reason="4">
<taxonomicName id="AFBF4D0F6656290233F3AA63FDC1196A" authority="(Womersley, 1952)" baseAuthorityName="Womersley" baseAuthorityYear="1952" box="[151,598,330,357]" class="Arachnida" family="Trombiculidae" genus="Helenicula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="signata">
<emphasis id="5ACBEA9E6656290233F3AA63FDC1196A" bold="true" box="[151,598,330,357]" pageId="86" pageNumber="87">
<emphasis id="5ACBEA9E6656290233F3AA63FEE5196B" bold="true" box="[151,370,330,357]" italics="true" pageId="86" pageNumber="87">Helenicula signata</emphasis>
(
<bibRefCitation id="0C2E4B7D6656290232E4AA63FDD9196A" author="Womersley, H." box="[384,590,330,356]" pageId="86" pageNumber="87" pagination="1 - 435" refId="ref110100" refString="Womersley, H. (1952) The scrub-typhus and scrub-itch mites (Trombiculidae, Acarina) of the Asiatic-Pacific region. Records of the South Australian Museum, 10, 1 - 435." type="journal article" year="1952">Womersley, 1952</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="20A565076656290233F3AA46FBBC1A33" pageId="86" pageNumber="87" type="reference_group">
<paragraph id="6800368C6656290233F3AA46FB9C1987" blockId="86.[151,1407,330,717]" box="[151,1035,366,393]" pageId="86" pageNumber="87">
<treatmentCitationGroup id="48AF11A26656290233F3AA46FB9C1987" box="[151,1035,366,393]" pageId="86" pageNumber="87">
<treatmentCitation id="E91E109D6656290233F3AA46FCFF1987" author="Womersley, H." box="[151,872,366,393]" page="174" pageId="86" pageNumber="87" year="1952">
<taxonomicName id="AFBF4D0F6656290233F3AA46FCFF1987" authority="Womersley 1952: 174" authorityName="Womersley" authorityPageNumber="174" authorityYear="1952" box="[151,872,366,393]" class="Arachnida" family="Trombiculidae" genus="Schoengastia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="signata" subGenus="Ascoschoengastia">
<emphasis id="5ACBEA9E6656290233F3AA46FDFF1986" box="[151,616,367,393]" italics="true" pageId="86" pageNumber="87">Schoengastia (Ascoschoengastia) signata</emphasis>
<bibRefCitation id="0C2E4B7D66562902310BAA46FCFF1987" author="Womersley, H." box="[623,872,366,393]" pageId="86" pageNumber="87" pagination="1 - 435" refId="ref110100" refString="Womersley, H. (1952) The scrub-typhus and scrub-itch mites (Trombiculidae, Acarina) of the Asiatic-Pacific region. Records of the South Australian Museum, 10, 1 - 435." type="journal article" year="1952">Womersley 1952: 174</bibRefCitation>
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, pl. 41 (A–D).
</treatmentCitationGroup>
</paragraph>
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<treatmentCitationGroup id="48AF11A26656290233F3AABAFD1B19A3" box="[151,652,402,429]" pageId="86" pageNumber="87">
<treatmentCitation id="E91E109D6656290233F3AABAFD1019A3" author="Radford, C. D." box="[151,647,402,429]" page="266" pageId="86" pageNumber="87" year="1954">
<taxonomicName id="AFBF4D0F6656290233F3AABAFD1019A3" authority=": Radford 1954: 266" authorityName="Radford" authorityPageNumber="266" authorityYear="1954" box="[151,647,402,429]" class="Arachnida" family="Trombiculidae" genus="Euschoengastia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="signata">
<emphasis id="5ACBEA9E6656290233F3AABAFE3519A2" box="[151,418,403,429]" italics="true" pageId="86" pageNumber="87">Euschoengastia signata</emphasis>
: 
<bibRefCitation id="0C2E4B7D6656290232D5AABAFD1019A3" author="Radford, C. D." box="[433,647,402,429]" pageId="86" pageNumber="87" pagination="247 - 276" refId="ref104337" refString="Radford, C. D. (1954) The larval genera and species of &quot; harvest mites &quot; (Acarina: Trombiculidae). Parasitology, 44, 247 - 276. https: // doi. org / 10.1017 / S 0031182000018898" type="journal article" year="1954">Radford 1954: 266</bibRefCitation>
</taxonomicName>
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.
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</paragraph>
<paragraph id="6800368C6656290233F3AA9EFC9619DF" blockId="86.[151,1407,330,717]" box="[151,769,438,465]" pageId="86" pageNumber="87">
<treatmentCitationGroup id="48AF11A26656290233F3AA9EFC9619DF" box="[151,769,438,465]" pageId="86" pageNumber="87">
<treatmentCitation id="E91E109D6656290233F3AA9EFD6A19DF" author="Audy, J. R." box="[151,765,438,465]" page="153" pageId="86" pageNumber="87" year="1954">
<taxonomicName id="AFBF4D0F6656290233F3AA9EFD6A19DF" authority=": Audy 1954: 153" authorityName="Audy" authorityPageNumber="153" authorityYear="1954" box="[151,765,438,465]" class="Arachnida" family="Trombiculidae" genus="Euschoengastia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="signata" subGenus="Helenicula">
<emphasis id="5ACBEA9E6656290233F3AA9EFDA119DE" box="[151,566,439,465]" italics="true" pageId="86" pageNumber="87">Euschoengastia (Helenicula) signata</emphasis>
: 
<bibRefCitation id="0C2E4B7D665629023120AA9EFD6A19DF" author="Audy, J. R." box="[580,765,438,465]" pageId="86" pageNumber="87" pagination="123 - 170" refId="ref95724" refString="Audy, J. R. (1954) Malaysian Parasites-IX. Notes on the taxonomy of trombiculid mites, with description of a new subgenus. Studies from the Institute for Medical Research, Federation of Malaya, 26, 123 - 170. https: // doi. org / 10.5962 / bhl. title. 119064" type="journal article" year="1954">Audy 1954: 153</bibRefCitation>
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.
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</paragraph>
<paragraph id="6800368C6656290233F3AAF2FBB419FB" blockId="86.[151,1407,330,717]" box="[151,1059,474,501]" pageId="86" pageNumber="87">
<treatmentCitationGroup id="48AF11A26656290233F3AAF2FBB419FB" box="[151,1059,474,501]" pageId="86" pageNumber="87">
<treatmentCitation id="E91E109D6656290233F3AAF2FC9B19FB" author="Chen, H. T. &amp; Hsu, P. K." box="[151,780,474,501]" page="126" pageId="86" pageNumber="87" year="1955">
<taxonomicName id="AFBF4D0F6656290233F3AAF2FC9B19FB" authority=": Chen &amp; Hsu 1955: 126" authorityName="Chen &amp; Hsu" authorityPageNumber="126" authorityYear="1955" box="[151,780,474,501]" class="Arachnida" family="Trombiculidae" genus="Globularoschoengastia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="signata">
<emphasis id="5ACBEA9E6656290233F3AAF2FE6119FA" box="[151,502,475,501]" italics="true" pageId="86" pageNumber="87">Globularoschoengastia signata</emphasis>
: 
<bibRefCitation id="0C2E4B7D665629023161AAF2FC9B19FB" author="Chen, H. T. &amp; Hsu, P. K." box="[517,780,474,501]" pageId="86" pageNumber="87" pagination="101 - 146" refId="ref97481" refString="Chen, H. T. &amp; Hsu, P. K. (1955) Report on 12 species and varieties of chiggers in China, including six new species and two new varieties, and the creation of a new genus. Acta Zoologica Sinica, 7, 101 - 146. [in both Chinese and English]" type="journal article" year="1955">Chen &amp; Hsu 1955: 126</bibRefCitation>
</taxonomicName>
</treatmentCitation>
; 
<treatmentCitation id="E91E109D66562902307CAAF2FBB719FB" author="Chen, H. T. &amp; Hsu, P. K." box="[792,1056,474,501]" page="250" pageId="86" pageNumber="87" year="1957">
<bibRefCitation id="0C2E4B7D66562902307CAAF2FBB719FB" author="Chen, H. T. &amp; Hsu, P. K." box="[792,1056,474,501]" pageId="86" pageNumber="87" pagination="379 - 404" refId="ref97542" refString="Chen, H. T. &amp; Hsu, P. K. (1957) Scrub mites from mammals and birds including the description of three new species. Acta Zoologica Sinica, 9, 379 - 404. [in Chinese]" type="journal article" year="1957">Hsu &amp; Chen 1957: 250</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="6800368C6656290233F3AAD6FBBC1A33" blockId="86.[151,1407,330,717]" pageId="86" pageNumber="87">
<treatmentCitationGroup id="48AF11A26656290233F3AAD6FBBC1A33" pageId="86" pageNumber="87">
<taxonomicName id="AFBF4D0F6656290233F3AAD6FDA51A17" authority=": Audy 1957: 266" authorityName="Audy" authorityPageNumber="266" authorityYear="1957" box="[151,562,510,537]" class="Arachnida" family="Trombiculidae" genus="Helenicula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="signata">
<emphasis id="5ACBEA9E6656290233F3AAD6FEFC1A16" box="[151,363,511,537]" italics="true" pageId="86" pageNumber="87">Helenicula signata</emphasis>
: Audy 1957: 266
</taxonomicName>
; 
<treatmentCitation id="E91E109D665629023159AAD6FC001A17" author="Womersley, H. &amp; Audy, J. R." box="[573,919,510,537]" page="281" pageId="86" pageNumber="87" year="1957">
<bibRefCitation id="0C2E4B7D665629023159AAD6FC001A17" author="Womersley, H. &amp; Audy, J. R." box="[573,919,510,537]" pageId="86" pageNumber="87" pagination="231 - 296" refId="ref110229" refString="Womersley, H. &amp; Audy, J. R. (1957) Malaysian Parasites-XXVII. The Trombiculidae (Acarina) of the Asiatic-Pacific Region: A revised and annotated list of the species in Womersley, 1952, with descriptions of larvae and nymphs. Studies from the Institute for Medical Research, Federation of Malaya, 28, 231 - 296." type="journal article" year="1957">Womersley &amp; Audy 1957: 281</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="E91E109D6656290230C7AAD6FA9B1A17" author="Nadchatram, M. &amp; Traub, R." box="[931,1292,510,537]" page="563" pageId="86" pageNumber="87" year="1971">
<bibRefCitation id="0C2E4B7D6656290230C7AAD6FA9B1A17" author="Nadchatram, M. &amp; Traub, R." box="[931,1292,510,537]" pageId="86" pageNumber="87" pagination="562 - 597" refId="ref103136" refString="Nadchatram, M. &amp; Traub, R. (1971) Chiggers of the genus Helenicula of the Old World including descriptions of 9 new species (Acarina: Prostigmata, Trombiculidae). Journal of Medical Entomology, 8, 562 - 597. https: // doi. org / 10.1093 / jmedent / 8.5.562" type="journal article" year="1971">Nadchatram &amp; Traub 1971: 563</bibRefCitation>
</treatmentCitation>
, figs. 1–7. 
<treatmentCitation id="E91E109D6656290233F3A90AFC0D1A33" author="Vercammen-Grandjean, P. H." box="[151,922,546,573]" page="112" pageId="86" pageNumber="87" year="1965">
<taxonomicName id="AFBF4D0F6656290233F3A90AFC0D1A33" authority=": Vercammen-Grandjean 1965 b: 112" authorityName="Vercammen-Grandjean" authorityPageNumber="112" authorityYear="1965" box="[151,922,546,573]" class="Arachnida" family="Trombiculidae" genus="Helenicula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="signata" subGenus="Helenicula">
<emphasis id="5ACBEA9E6656290233F3A90AFD971A32" box="[151,512,547,573]" italics="true" pageId="86" pageNumber="87">Helenicula (Helenicula) signata</emphasis>
: 
<bibRefCitation id="0C2E4B7D66562902316AA90AFC0D1A33" author="Vercammen-Grandjean, P. H." box="[526,922,546,573]" pageId="86" pageNumber="87" refId="ref107633" refString="Vercammen-Grandjean, P. H. (1965 b) Trombiculinae of the world. Synopsis with generic, subgeneric, and group diagnoses (Acarina, Trombiculidae). George Williams Hooper Foundation, University of California, San Francisco, California, 191 pp." type="book" year="1965">Vercammen-Grandjean 1965b: 112</bibRefCitation>
</taxonomicName>
</treatmentCitation>
; 
<treatmentCitation id="E91E109D6656290230C1A90AFBB01A33" author="Vercammen-Grandjean, P. H." box="[933,1063,547,573]" page="101" pageId="86" pageNumber="87" year="1968">
<bibRefCitation id="0C2E4B7D6656290230C1A90AFBB01A33" author="Vercammen-Grandjean, P. H." box="[933,1063,547,573]" pageId="86" pageNumber="87" refId="ref107767" refString="Vercammen-Grandjean, P. H. (1968 b) The chigger mites of the Far East (Acarina: Trombiculidae &amp; Leeuwenhoekiidae). U. S. Army Medical Research and Development Command, Washington, D. C., 135 pp." type="book" year="1968">1968b: 101</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="20A565076656290233F3A96EFE621AC3" pageId="86" pageNumber="87" type="materials_examined">
<paragraph id="6800368C6656290233F3A96EFE2F1A6F" blockId="86.[151,1407,330,717]" box="[151,440,583,609]" pageId="86" pageNumber="87">
<typeStatus id="B704882E6656290233F3A96EFF581A6F" box="[151,207,583,609]" pageId="86" pageNumber="87">Type</typeStatus>
deposition. No data.
</paragraph>
<paragraph id="6800368C6656290233F3A942FC201A8B" blockId="86.[151,1407,330,717]" box="[151,951,619,645]" pageId="86" pageNumber="87">
<materialsCitation id="D8D73CD16656290233F3A942FC241A8B" ID-GBIF-Occurrence="3026169719" box="[151,947,619,645]" country="Brunei" location="Labuan" pageId="86" pageNumber="87" specimenCount="1" stateProvince="Labuan" typeStatus="holotype">
<typeStatus id="B704882E6656290233F3A942FF581A8B" box="[151,207,619,645]" pageId="86" pageNumber="87">Type</typeStatus>
data. Free larva collected on boots, 
<collectingCountry id="10A8761C665629023100A942FD271A8B" box="[612,688,619,645]" name="Brunei" pageId="86" pageNumber="87">Brunei</collectingCountry>
and 
<collectingCountry id="10A8761C665629023183A942FCC71A8B" box="[743,848,619,645]" name="Malaysia" pageId="86" pageNumber="87">Malaysia</collectingCountry>
, 
<collectingRegion id="AA7BF86E66562902303FA942FC241A8B" box="[859,947,619,645]" country="Malaysia" name="Labuan" pageId="86" pageNumber="87">Labuan</collectingRegion>
</materialsCitation>
.
</paragraph>
<paragraph id="6800368C6656290233F3A9A6FDF31AA7" blockId="86.[151,1407,330,717]" box="[151,612,655,681]" pageId="86" pageNumber="87">
Hosts. 
<taxonomicName id="AFBF4D0F665629023381A9A6FE0B1AA7" box="[229,412,655,681]" kingdom="Plantae" order="Scandentia" pageId="86" pageNumber="87" rank="order">SCANDENTIA</taxonomicName>
: 
<taxonomicName id="AFBF4D0F6656290232C3A9A6FDCA1AA6" authorityName="Thomas" authorityYear="1892" box="[423,605,655,680]" class="Mammalia" family="Tupaiidae" genus="Tupaia" kingdom="Animalia" order="Scandentia" pageId="86" pageNumber="87" phylum="Chordata" rank="species" species="montana">
<emphasis id="5ACBEA9E6656290232C3A9A6FDCA1AA6" box="[423,605,655,680]" italics="true" pageId="86" pageNumber="87">Tupaia montana</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="6800368C6656290233F3A99AFE621AC3" blockId="86.[151,1407,330,717]" box="[151,501,691,717]" pageId="86" pageNumber="87">
Distribution. 
<collectingCountry id="10A8761C665629023249A99AFEEB1AC3" box="[301,380,691,717]" name="Brunei" pageId="86" pageNumber="87">Brunei</collectingCountry>
, 
<collectingCountry id="10A8761C6656290232E3A99AFE661AC3" box="[391,497,691,717]" name="Malaysia" pageId="86" pageNumber="87">Malaysia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
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