Review of the family Coccidae (Hemiptera: Coccomorpha) in Laos Author Choi, Jinyeong Author Soysouvanh, Pheophanh Author Lee, Seunghwan Author Hong, Ki-Jeong text Zootaxa 2018 2018-08-17 4460 1 1 62 journal article 29001 10.11646/zootaXa.4460.1.1 84973bd6-9d66-4172-8f50-2a482edccf2a 1175-5326 1459506 DB841017-698F-4D44-A633-461D350DC984 Genus Megapulvinaria Yang, 1982 Type species: Pulvinaria maxima Green, 1904a , by original designation. Diagnosis. Dorsum with anal plates each with 1 spinose discal seta, and 2 spinose and/or truncate setae on inner margin and apex ( Figs 35D , 36G ); tubular ducts present or absent ( Fig. 36F ); duct tubercles absent. Marginal setae stout and truncate, with 2 types (one long and slender; the other shorter or same length and thicker) ( Figs 35C , 36B ). Stigmatic clefts shallow to deep, each containing 3–12 stigmatic spines ( Fig. 36C ). Venter with tubular ducts of 3 types ( type I: each with a broad inner ductule, present on medial area of head, thorax and anterior abdomen; type II: each with a narrow inner ductule, present on medial and inner submarginal area of abdomen; type III: each with a filamentous inner ductule, present in submarginal areas) ( Fig. 36J ); antenna 7 to 9 segmented ( Fig. 36M ); legs each with tibio-tarsal articulatory scleroses ( Fig. 36I ) ( Hodgson 1994 ; Wang & Feng 2012b ). Remarks. Megapulvinaria contains 5 described species that are mainly distributed in the Oriental and Australian regions ( García Morales et al . 2016 ). The genus is related to Pulvinarisca , but is easily differentiated from the latter by possessing 2 types of marginal setae, and each anal plate having 1 discal seta and 2 spinose and/ or truncate setae ( Hodgson 1994 ); in contrast, Pulvinarisca has only 1 type of marginal seta and each anal plate has 3 spinose setae only ( Hodgson 1994 ; Wang & Feng 2012b ).