On the species-group taxa of Taiwanese social wasps (Hymenoptera: Vespidae) described and / or treated by J. Sonan
Author
Kojima, Jun-Ichi
Author
Saito, Fuki
Author
Nguyen, Lien Thi Phuong
text
Zootaxa
2011
2920
42
64
journal article
46640
10.5281/zenodo.201805
f9385d5f-2207-403f-b7e9-4bd5440581be
1175-5326
201805
Vespa formosana
Sonan 1927
[
Vespa affinis
Linnaeus 1764
]
Sonan (1927)
did not designate the
holotype
when he described this nominal taxon based on queen(s) [body length
28 mm
; wing span
50 mm
; fore wing length
21 mm
], worker(s) [body length
22 mm
; wing span
35 mm
; fore wing length
16 mm
] and male(s) [body length
21 mm
; wing span
41 mm
; fore wing length
18 mm
] from “Taihoku, Kagi, Takezaki, Nanto, Daimokko, Maruyama”.
Starr (1992: 102)
mentioned “
types
examined:
1 queen
,
1 worker
, 1 3”, which does not constitute
lectotype
designation.
Type
material.
In
TARI
, we located only the following specimens that could be from the
type
series of
Vespa formosana
.
LECTOTYPE
(designated here): one queen (body length
28 mm
; wing span
46 mm
; fore wing length
20 mm
), with labels of
Type
label, “Taihoku /
3.VII 1926
/ Col. J. Sonan”, “
Vespa formosana
/ Sonan”, and “161”.
PARALECTOTYPES
: one worker (body length
21 mm
; wing span
33 mm
; fore wing length
16 mm
), with
Type
label, “Ƥ
type
”, “Kagi /
14 VI 1926
/ Toyota”, “
Vespa formosana
/ Sonan” and “162”; and 1 3 (body length
19 mm
; fore wing length
18 mm
), with
Type
label, “
type
3”, “
Vespa formosana
/ Sonan”, and “163” but without any collection data label. These specimens could be those that
Starr (1992)
refered to as “
types
examined”.
Sonan (1929)
listed the following specimens that he examined:
1 worker
, Dakusui (Ratô),
4.XI.1928
(J. Sonan);
1 worker
, Fukô,
1.VII.1925
(J. Sonan);
1 worker
*, Kagi,
25.VI.1926
(K. Toyota);
2 workers
*, Kagi,
10.VI.1926
(K. Toyota);
1 worker
*, Kagi,
XI.1923
(M. Kato);
1 queen
*, Kagi,
24.VII.1924
(K. Toyota);
1 queen
*, Nanto,
IV.1910
(G. Toba);
1 queen
*, Nanto,
IV.1910
(T. Shiraki);
1 worker
, Sozan,
29.IX.1926
(J. Sonan);
1 worker
*, Taihoku,
27.VIII.1908
(I. Nitobe);
1 worker
*, Taihoku,
28.II.1913
(S. Inamura);
1 queen
*, Taihoku,
XI.1912
(I. Nitobe);
1 queen
, Taihoku,
23.V.1928
(J. Sonan);
1 queen
*, Taihoku,
VII.1925
(J. Sonan);
1 queen
*, Taihoku,
5.VI.1908
(I. Nitobe);
1 queen
*, Taihoku, 3.
VII.1
[9]26 (J. Sonan);
1 queen
*, Taihoku,
XI.1912
(I. Nitobe);
2 workers
*, Taihoku,
30.VIII.1925
(J. Sonan);
1 worker
*, Takezaki,
15.IX.1926
(J. Sonan). Among these specimens those with asterisks had been collected before year 1927 when the original description of
Vespa formosana
was published, and at the localities mentioned in the original description; they could be examined by Sonan when he prepared the original description. However,
Sonan (1927)
did not mention any variation in color pattern or morphology, and thus we conclude that the original description was based on the three specimens (one queen, one worker and one male) with the
Type
labels. The queen specimen from “Taihoku” that has a label of specimen number “161” is here desiganated as the
lectotype
of
Vespa formosana
Sonan 1927
. The worker from “Kagi” (specimen number “162”) and the male with specimen number “163” are
paralectotypes
.
Bequaert (1936: 345)
, on the strength of the original descriptions and specimens from “Takao” and “Hu Sia” [unrecognized locality], synonymized this nominal taxon under
Vespa affinis
(Linnaeus)
. Later he (
Bequaert 1939: 42
) confirmed his own synonymy, mentioning “Dr. K. Yasumatsu sent me recently a specimen of what is called
V.
formosana
by Japanese entomologists.” However, Bequaert's synonymy has long been overlooked. Van der
Vecht (1957
,
1959
) did not refer to this synonymy,
Kuo (1984)
and
Kuo & Yeh (1987)
used
V. f o r m o s a n a
to describe ecological aspects of this species, and finally
Starr (1992)
referred to Sonan's
types
and recognized the synonymy of
V. formosana
with
V. affinis
but did not refer to Bequaert's synonymy. The
lectotype
(and also
paralectotypes
) of
Vespa formosana
here designated belongs to
V. affinis
and the synonymy of
Bequaert (1936)
has been confirmed.