Late Cretaceous dinosaurs from the Denver Basin, Colorado
Author
Kenneth Carpenter
Department of Earth Sciences, Denver Museum of Nature & Science, 2001 Colorado Boulevard, Denver, CO 80205, U. S. A.
Author
D. Bruce Young
Department of Earth Sciences, Denver Museum of Nature & Science, 2001 Colorado Boulevard, Denver, CO 80205, U. S. A.
text
Rocky Mountain Geology
2002
37
237
254
journal article
10.2113/gsrocky.37.2.237
41bd81a8-8d16-41cb-950f-8fab1660f845
3943081
Ceratopsid
genus and species indet.
Figures 3
,
8
, and
17
Polyonax mortuarius
Cope, 1874a
Bison
alticornis
Marsh, 1887
Ceratops alticornis
Marsh, 1889
TYiceratops alticornis
Hatcher, 1907
Ceratops montanus
Marsh, 1888
TYiceratops galeus
Marsh, 1889
Material.
—AMNH 3950 dorsal centra, caudal centra, rib fragment, fibula fragment (holotype of
Polyonax mortuaris-
,
Laramie Formation, East Bijou Creek); DMNH 2823 horn core, vertebra fragment (Denver Formation, Loc. 19); DMNH 25914 braincase (Denver Formation, Loc. 18); DMNH 25915 braincase, horn core fragment, frill fragment, dentary fragment with teeth (Denver Formation, Loc. 20); DMNH 27743 tooth (Denver Formation, Loc. 20); DMNH 27744 tooth (Denver Formation, Loc. 20); DMNH 29951 tooth (Denver Formation, Loc. 20); DMNH 33301 tooth (Denver Formation, Loc. 20); DMNH 33351 horn core, squamosal fragment (Laramie Formation, Loc. 8); DMNH 34291 left mandible (Denver Formation, Loc. 20); DMNH 34391 dentary, parietal fragment with base of horn core, left squamosal fragment (Denver Formation, Loc. 20); DMNH 34392 section of horn (Denver Formation, Loc. 20); DMNH 34595 radius? fragment (Denver Formation, Loc. 21); DMNH 44392 braincase (Denver Formation, Loc. 18); UCM 68914 right articular, posterior dorsal, neural spine, rih fragments, left humeral shaft, right partial ilium, right ischium, distal end of left ischium, left pubis (Denver Formation, Loc. 31); UCM 13746 horn core (Laramie Formation, Loc. 3); UCM 13749 horn core (Laramie Formation, Loc. 3); UCM 36398 humeral shaft, femoral shaft, rib fragments (Denver
Formation
, Loc. 22); UCM 42328 teeth (Laramie Formation, Loc. 1); UCM 43699 braincase (Laramie Formation, Loc. 2); UCM 47634 dentary fragment (Denver Formation, Loc. 27); UCM 47635 tooth (Denver Formation, Loc. 27); USNM 2410 nasal horn core (holotype of
Triceratops galeus-
,
Arapahoe Formation, Loc. 11); USNM 4739 pair of frontals and horn cores (holotype of
“Bison"
alticornis
)
Denver Formation, Loc. 29); and USNM uncataloged (Arapahoe Formation, Locs. 21, 22).
Description and
discussion. —The hulk of the ceratopsian remains from the Denver Basin cannot he identified below family level. Most of the specimens indicate large, hence adult individuals. For example, the nearly complete dentary, DMNH 34391, is 52 cm in length and has 24 alveoli. A fragment of the predentary is present and is slightly curved, measuring 9.5 cm in length.
The material
Cope (1874a)
named
Polyonax mortuaris
from the eastern part of the Denver Basin is badly abraded and fragmented. In his review of the ceratopsians,
Hatcher (1907)
concluded that none of the material can he assigned to any of the known ceratopsian taxa, a point with which we concur. Marsh (1888) tentatively referred some bones from Colorado, which
Lull (1907)
identified as from the Arapahoe Formation as belonging to
Ceratops montanus
(these are uncataloged USNM specimens).
Lull (1907)
noted that the referral is improbable because of the vast age difference between the Judith River and the Arapahoe Formations. He further noted that the material is too fragmentary for positive identification.
Figure
17.
Restored pelvis of the ceratopsian shown in Figure 8 in right lateral view (pubis reversed). Scale bar = 10 cm.
Two holotypes of Marsh are also among the indeterminate ceratopsians. Marsh (1889) named
Triceratops
galeus
for a nasal horn core collected from the Arapahoe Formation (Loc. 13).
Hatcher (1907)
noted that the specimen closely resembles that of the
Torosaurus
,
but Lull (1933) noted that it also resembles that of a
Triceratops
skull, AMNH 5116 (compare
Hatcher, 1907
, fig. 111 with
Forster
, 1996
, fig. 4A). He therefore considered
T. galeus
a nomen dubium, a point with which we concur. Another holotype is the famous pair of brow horn cores originally named by Marsh as
Bison alticornis
,
but later referred by him (Marsh, 1889) to
Ceratops
alticornis
. More recently, the specimen has been referred to
Triceratops
(
Hatcher, 1907
;
Lull, 1907
; Lull, 1933; Ostrom and Welnhofer, 1986), although
Forster
(1996)
noted that the horns cannot be assigned to
Triceratops
with certainty, a conclusion independently reached here. The horns most certainly could belong to
Diceratops
or
Torosaurus
as readily as to
Triceratops
.
A partial ceratopsian skeleton was collected from the flanks of South Table Mountain (
Fig. 8
) by the Colorado School of Mines. The specimen has since been transferred to the University of Colorado Museum. The specimen includes pelvic material, which allows a composite restoration to be made (
Fig. 17
). The pelvis is rather small compared to those in the Smithsonian collections from the Lance Formation.