The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera Author Harzhauser, Mathias 0000-0002-4471-6655 mathias.harzhauser@nhm-wien.ac.at Author Landau, Bernard 0000-0002-7768-8494 bernardmlandau@gmail.com Author Janssen, Ronald 0000-0002-4471-6655 mathias.harzhauser@nhm-wien.ac.at text Zootaxa 2022 2022-03-30 5123 1 1 172 http://dx.doi.org/10.11646/zootaxa.5123.1.1 journal article 10.11646/zootaxa.5123.1.1 1175-5326 6399827 036F6B4D-CDCC-4CD7-A914-9A1D8C7A097A Striopusionella szekelyhidiae ( Kovács & Vicián, 2021 ) nov. comb. Figs 42B 1 –B 2 , 4 Ab, 5, 7 * Clavatula szekelyhidiae n. sp. — Kovács & Vicián 2021: 145 , pl. 3, figs 7–10. Type material. Holotype : HNHM , PAL 2021.19 .1, SL: 65.0 mm, MD : 25.6 mm , Letkés ( Hungary ), illustrated in Kovács & Vicián (2021 , pl. 3, figs 9–10); figs 42B 1 –B 3 , 4 Ab . Paratype : NHMW 2021 /0002/0002, SL: 53.5 mm , MD : 19.4 mm , Letkés ( Hungary ), illustrated in Kovács & Vicián (2021 , pl. 3, figs 7–8) . Revised description. Shell moderately large, solid, moderately slender fusiform with high conical spire; apical angle ~35°. Protoconch not preserved. Teleoconch of eleven whorls. Early teleoconch whorls with tripartite sculpture, weakly concave with two beaded adsutural cords, separated by wide concavity with weak central cord of delicate beads. Later teleoconch whorls broadly and weakly concave, subsutural collar narrow, weak, bearing small, wide-spaced tubercles, subobsolete in some specimens, suprasutural tubercles almost completely obscured by succeeding whorl. Entire shell covered by raised, narrow convex spiral cords, separated by interspaces of roughly equal width. Suture narrowly impressed, finely undulating. Last whorl 60% of total height. Subsutural collar very poorly delimited. Subsutural ramp broad, weakly concave, delimited and angled by narrow weakly tuberculose shoulder cord. Straight sided below, strengthened by very weakly tuberculose peribasal cord. Base moderately constricted, weakly concave in profile. Sinuous, poorly delimited axial folds starting on the abapical half of ramp and extending to base. Siphonal fasciole poorly delimited, strongly rounded and twisted, forming wide, deep pseudoumbilicus in fully adult specimens. Sculpture below shoulder cord of narrow irregularly beaded cords of roughly alternate strength. Aperture moderately wide, ovoid pyriform. Outer lip not thickened, smooth within. Anal sinus very wide, very shallow with apex mid-ramp. Siphonal canal short, narrow. Columella weakly excavated, smooth. Columellar and parietal callus strongly thickened, sharply delimited, moderately broad. Discussion. We are not aware of comparable species in the Paratethyan fauna. ‘ Clavatula ’ occidentalis Peyrot, 1931 , from the Langhian of Saubrigues ( France ), displays a similar sculpture, but differs from the Paratethyan species in its shorter spire, longer siphonal canal and more concave spire whorls. Its anal canal is much deeper and therefore a closer relation with Striopusionella szekelyhidiae ( Kovács & Vicián, 2021 ) is unlikely (see Peyrot 1931 , pl. 5, figs 10, 18). Similarly, ‘ Clavatula ’ saubrigniana ( Grateloup, 1832 ), from the Langhian of Saubrigues ( France ), is similarly slender in outline, but has a deep anal sinus (see Peyrot, 1931 , pl. 5, figs 21, 26, 27). Paleoenvironment. The occurrence at Letkés ( Hungary ) suggests a preference for inner neritic environments in the vicinity of corals ( Kovács & Vicián 2013 ). Distribution in Central Paratethys. Badenian (middle Miocene): Pannonian Basin : Letkés ( Hungary ) ( Kovács & Vicián 2021 ).