Lectotypes of Erebia kefersteinii (Eversmann, 1851), Clossiana selenis (Eversmann, 1837), Melitaea arcesia Bremer, 1861 and holotype of Melitaea baikalensis Bremer, 1861 (Lepidoptera: Nymphalidae)
Author
Churkin, Sergei V.
0000-0001-9195-0340
Nosovikhinskoye shosse, 3, Reutov, 143965, Russia. E-mail: serghelios 16 @ gmail. com, https: // orcid. org / 0000 - 0001 - 9195 - 0340
Author
Yakovlev, Roman V.
0000-0001-9512-8709
Altai State University, Lenina pr. 61, Barnaul, 656049, Russia. E-mail: yakovlev _ asu @ mail. ru, https: // orcid. org / 0000 - 0001 - 9512 - 8709 & Tomsk State University, Lenina pr. 36, 634050 Tomsk, Russia.
Author
Lvovsky, Alexandr L.
0000-0002-7170-8773
Zoological Institute, Universitetskaya nab., 1, Sankt-Petersburg, 199034, Russia. E-mail: Alexander. Lvovsky @ zin. ru, https: // orcid. org / 0000 - 0002 - 7170 - 8773
text
Ecologica Montenegrina
2022
2022-08-30
55
17
25
http://dx.doi.org/10.37828/em.2022.55.2
journal article
10.37828/em.2022.55.2
2336-9744
13241051
Erebia kefersteinii
(
Eversmann, 1851
)
Bull. Soc. imp. Nat.
Moscou
24 (2): 610–611
Originally published as “
Hipparchia (Erebia) kefersteinii
“
Type locality after Eversmann: “…Siberie orientale.”
Two syntypes
, male and female, were found in the former Eversmann collection in
ZISP
. At least 4 more subspecies are known, thus, to preserve the stability of zoological nomenclature, it is important to have the name-bearing type of the taxon fixed by the
lectotype
designation
.
Figures 1−15.
Erebia kefersteinii
Eversmann, 1851
, adult specimens and labels: 1.
E. kefersteinii
, lectotype, female, upperside (ZISP); 2.
E. kefersteinii
, lectotype, female, underside (ZISP); 3.
E. kefersteinii
, lectotype, labels; 4.
E. kefersteinii
, paralectotype, female, upperside (ZISP); 5.
E. kefersteinii
, paralectotype, female, underside (ZISP); 6.
E. kefersteinii
, paralectotype, labels; 7.
E. kefersteinii
, male, Kultuk village, road to Khamar-Daban, upperside (ZISP); 8.
E. kefersteinii
, male, labels; 9.
E. kefersteinii amika
,
paratype, male, upperside (coll. S. Churkin, Reutov); 10.
E. kefersteinii amika
,
paratype, male, underside (coll. S. Churkin, Reutov); 11.
E. kefersteinii amika
,
paratype, female, upperside (coll. S. Churkin, Reutov); 12.
E. kefersteinii amika
,
paratype, female, underside (coll. S. Churkin, Reutov); 13.
E. kefersteinii amika
,
paratype, female, upperside (coll. S. Churkin, Reutov); 14.
E. kefersteinii amika
,
paratype, female, underside (coll. S. Churkin, Reutov); 15.
E. kefersteinii amika
,
labels.
Lectotype
(
Figs 1–3
): female, forewing length
18 mm
, with three original labels:
- printed label "coll Eversmann" [Eversmann’s collection];
- small handwritten rectangular label "Irkutzk" [
Irkutsk
] (certainly, Eversmann’s hand);
- small rectangular label with two words written illegibly (we spent a lot of time searching different old maps of the region trying to understand the name of the locality and even asking the help of the
Irkutsk
History Museum but all efforts were without any success).
We add the red label “
LECTOTYPUS
/
Hipparchia (Erebia)
/
kefersteinii
Eversmann
,/ 1851 S.CHURKIN et al. des.“
The specimen has no antennae (except the part of the right one), the coloration is fully similar to the description, full series of the blackish dots are situated on the fore- and hindwings.
Paralectotype
(
Figs 4–6
): male, forewing length 18.2 mm, all labels are the same as under the
lectotype
. The specimen is not fresh and partly damaged (left forewing and both hindwings with some chips).
We add the red label “
PARALECTOTYPUS
/
Hipparchia (Erebia)
/
kefersteinii
Eversmann
,/ 1851 S.CHURKIN et al. des.“
Type locality
Unfortunately, the true name of the collecting locality is written illegibly. Undoubtedly, this place is located somewhere on the road to Baikal, only one available road here at that times.
In addition, we found in
ZISP
collection a small series of the specimens from
Erschoff
collection, also very old specimens collected in 1878, some are with the reddish label “
Irkutsk
”.
Some
butterflies have additional label “Kultuk” and only one has the four-fold paper with the explanation: «село Култук, дорога на Хамар- Дабан» (Kultuk village, road to Khamar-Daban”) (
Figs 7–8
).
All
these specimens are not the
types
, but look very similar to those, and it is very possible that it were collected according to the information where the first known
kefersteinii
were caught. Worth to note that Kultuk is situated at the mentioned Baikal road, it was well-known collecting place
.
Thus, the
type
locality is not far from Kultuk vicinity,
NW
Baikal Lake
,
Irkutsk
reg
.
Status and notes
The species demonstrates obvious geographical clines and forms, which are masked by individual variability. The butterflies from the
Altai
have clear reddish hue and short median band. The butterflies from the Khamar-Daban and the
East Sayan
have broader median band and clearly yellowish with reduced reddish hue (i.e. belong to the nominate taxon) while the
West Sayan
and
Tuva
populates by the specimens with all variants of mixed colouration. The representatives of
ssp.
amica
Churkin, 1999
from the most eastern point of the distribution area – the Barguzinsky Range – are characterized by reddish coloration as the specimens from the
Altai
, but usually have broad band as the specimens from the Khamar-Daban and the
East Sayan
.
We are able to figure here the good colored photo of the
paratypes
of this taxon (
Figs 9–15
,
one male
and
two females
).
Two subspecies –
ssp.
kholsunica
Lukhtanov, 1990
(
type
locality – South
Altai
, Kholzun range) and
ssp.
otteni
Murzin & Sinayev, 2003
(
type
locality – Kuznetskyi Alatau) are characterised by the reduced black dots.
The simple idea that butterflies from high altitudes have reduced dots is wrong: the
type
series of
otteni
without the spots and the
type
series of
amica
with full number of spots were collected at the same altitudes.
In such a situation it is impossible to unite all known geographical forms under one subspecies name, but true system of the subspecies is needed in revision.