A world revision of the Pycnostigminae (Cynipoidea: Figitidae) with descriptions of seven new species Author Noort, Van text Zootaxa 2007 1392 1 30 journal article 10.5281/zenodo.175269 4ede1b61-2f74-4fdc-b50e-c3ea3ebd4c49 1175-5326 175269 Pycnostigmus rostratus Cameron ( Figs 1–2 ) Pycnostigmus rostratus Cameron 1905 : 20 –21. Type species by monotypy. Redescription. Holotype FEMALE. Head . Shagreened and substantially setose (frons, occiput, cheeks) ( Figs 1 E & 2E). Antenna entirely dark brown/black, scape 1.75x length of radicle, short appressed setae on all 16 flagellomeres, moniliform. Mesosoma . Lateral surface of pronotum densely rugulose and pitted; evenly covered in moderately long setae ( Figs 1 B & 2B). Mesopleuron striate except for small area immediately ventrad of mesopleural triangle, and an additional portion immediately dorsal of mesocoxa; dorsal and ventral margins of mesopleural triangle clearly visible, center of triangle setose ( Figs 1 B & 2B). Mesoscutum weakly pitted with small punctures and sparsely setose; cuticle between punctures shiny; notaulices complete, originating at anterior end of parascutal impression; posterior convergence point of notaulices with a small subquadrate, shiny portion of the mesoscutum indented ( Figs 1 C & 2C). Disk of scutellum strongly rugose and deeply pitted, setose along posterior margin; sides not marked by a distinct carina, curving ventrally ( Figs 1 C & 2C); scutellar ridge that separates scutellar fovea often extending to small spine along posterior margin of scutellum. Metapleural-propodeal complex . Entire metapleuron covered in long, thin setae ( Figs 1 B & 2B). Propodeum sparsely covered by long setae; cuticle deeply rugulose across entire propodeum. Nucha extended posteriorly, deeply striate ( Figs 1 A, 2A, 1C & 2C). Metasoma . Petiole about as long as deep, deeply striate. Syntergum (T2+T3) shiny and smooth; posterior portion of T-3-4 with a brush of setae ( Figs 1 A & 2A). Entire metasoma, in lateral view, about as long as deep. Wings . Pseudostigma weakly pigmented, marginal cell visible ( Figs 1 D & 2D). Legs. Femora and tibiae black; long setae present. Tarsomeres brown, covered in short, appressed setae ( Figs 1 A & 2A). MALE. As in female except: 18 antennal flagellomeres present; length of metasoma 1.67x depth. Diagnosis. Separated from P. mastersonae by the head and mesosoma being non-metallic; from P . fossilensis by having the disk of the scutellum completely rugose (center smooth in P . fossilensis ); from P . incognito by having the mesopleuron mostly striate (almost entirely smooth in P . incognito ); from P. hoerikwaggonensis by having the lateral aspect of the pronotum areolate (striate in P. hoerikwaggonensis ). Distribution. South Africa : Western Cape ( Fig. 13 ). Material examined. Holotype : [first label] Cape T, 9/87 [Cape Town, September, 1887], [second label] SAM-HYM-P002866, [third label, in Cameron’s handwriting] Pycnostigmus rostratus Cam. Type Cape Town (see inset Fig. 1 A for labels). The type is a female, in good condition (c.f. Fig. 1 ), having been remounted on a card by M. Sporrong ( Sweden ). Deposited in SAMC . Additional material. Paratype : SOUTH AFRICA . [first label] Type ( NHML type label with red ring), [second label] B.M. Type Hym. 7. 1., [third label, in Cameron’s handwriting] Pycnostigmus rostratus Cam. Type Cape Town, [fourth label] Cameron Coll. 1906–138. The type is in moderate condition, with the head plus mesosoma together and mounted separately from the metasoma. Deposited in NHML . Western Cape Province . Koeberg Nature Reserve, 33°37.622'S 18°24.259'E , 8 Aug–5 Sept 1997 , S van Noort, Malaise trap KO97-M07, West Coast Strandveld, ( 1 female ) [ SAMC ]; Koeberg Nature Reserve, 33°37.622'S 18°24.259'E , 8 Aug–5 Sept 1997 , S van Noort, Malaise trap KO97-M08, West Coast Strandveld, SAM-HYM-P024713 ( 1 female ) [ SAMC ]; West Coast Fossil Park ( 5.5 km 290o W Langebaanweg) 32o57.402’S 18o05.438’E , 4-11 Sept 2002 , S van Noort, Malaise trap, LW02-R5-M12, rehabilitated mine dump, SAM-HYM-P024398 (determined by F. Ronquist, 2005) ( 1 female ) [ SAMC ]; Jacobsbaai, N of Saldanha Bay, 32o58’S 17o54’E , P.T. O ., HG Robertson, West Coast Strandveld ex pitfall trap, SAM-HYM-P024396 ( 1 female ); West Coast Fossil Park, ( 3 km 270° W Langebaanweg), 32°58.445'S 18°07.236'E , 18–25 Sept 2002 , S van Noort, Malaise trap LW02-N3-M39, Sand Plain Fynbos ( 4 female , 2 male ) [ SAMC ]; Anyskop Farm, ( 5.5 km 290° W Langebaanweg) 32°57.394'S 18°05.324'E , 16–23 Oct 2002 , S van Noort, Malaise trap LW02-N7-M115, Sand Plain Fynbos ( 1 female , 1 male ) [ USNM ], West Coast Fossil Park, ( 3 km 270° W Langebaanweg) 32°58.445'S 18°07.236'E , 18–25 Sept 2002 , S van Noort, Malaise trap LW02-N3-M39, Sand Plain Fynbos ( 1 male ) [ SAMC ]; Anyskop Farm, ( 5.5 km 290° W Langebaanweg) 32°57.301'S 18°05.283'E , 18–25 Sept 2002 , S van Noort, Malaise trap LW02-N8-M48, Sand Plain Fynbos ( 2 females ) [ SAMC ]; Langberg Farm, ( 3 km 280° W Langebaanweg) 32°58.045'S 18°07.418'E , 18–25 Sept 2002 , S van Noort, Malaise trap LW02-N1-M35, Sand Plain Fynbos ( 1 male , 1 female ) [ CASC ]; Anyskop Farm, ( 5.5 km 290° W Langebaanweg) 32°57.394'S 18°05.324'E , 25.ix–2.x 2002 , S van Noort, Malaise trap LW02-N7-M64, Sand Plain Fynbos SAM-HYM- P0024414 ( 4 females , 2 males ) [ SAMC ]; Anyskop Farm, ( 5.5 km 290° W Langebaanweg) 32°57.394'S 18°05.324'E , 9–16 Oct 2002 , S van Noort, Malaise trap LW02-N7-M98, Sand Plain Fynbos ( 2 females ) [ SAMC ]; De Hoop Nature Reserve, 34°27'S 20°25'E 11.X.1994 , coastal fynbos, sweepnet, leg. Michael Söderlund ( 1 male ) [ ULMZ ]. FIGURE 1. Pycnostigmus rostratus Cameron , holotype specimen, female. A, habitus; inset, specimen labels. B, head and mesosoma, lateral view. C, head and mesosoma, dorsal view. D, left forewing. E, head, anterior view. FIGURE 2. Pycnostigmus rostratus Cameron , non-type specimen, female. A, habitus. B, head and mesosoma, lateral view. C, head and mesosoma, dorsal view. D, left forewing. E, head, anterior view. Image deposition. A MorphBank collection of images of P. rostratus can be accessed via http://morphbank.net/Show/?id= 110332 .