Further evidence of Cretaceous termitophily: Description of new termite hosts of the trichopseniine Cretotrichopsenius (Coleoptera: Staphylinidae), with emendations to the classification of lower termites (Isoptera) Author JIANG, RI-XIN Author ZHANG, HONG-RUI Author ELDREDGE, K. TARO Author SONG, XIAO-BIN Author LI, YAN-DA Author TIHELKA, ERIK Author HUANG, DI-YING Author WANG, SHUO Author ENGEL, MICHAEL S. Author CAI, CHEN-YANG text Palaeoentomology 2021 2021-08-31 4 4 374 389 http://dx.doi.org/10.11646/palaeoentomology.4.4.13 journal article 10.11646/palaeoentomology.4.4.13 6cd3d259-465d-4ab3-8a32-d57a812bb6e0 2624-2834 5507925 D1F5A853-32D1-470B-A609-D54F7B8C60CA Tanytermitalis philetaerus Engel & Cai , sp. nov. ( Figs 1B, D , 7 ) Holotype . Holotype alate, NIGP174707 . In mid- Cretaceous amber from Kachin Province , northern Myanmar . Deposited in the Nanjing Institute of Geology and Palaeontology , Chinese Academy of Sciences , Nanjing , China . Etymology. The specific epithet is taken from the Greek term ΦΙΛέταΙΡΟΣ ( philetairos ) [composed of ΦῘÌΛΟΣ ( phílos ), meaning, “beloved”, and ἑταῖΡΟΣ ( hetaîros ), meaning, “friend”), and refers to the relationship between these termites and their inquilines. Diagnosis. As for the genus ( vide supra ). Description. Imago: Total body length (as preserved) 5.28 mm ; length of head to apex of labrum 1.12 mm ; head width 0.68 mm ; compound eye diameter 0.22 mm ; distance from compound eye to posterior margin 0.36 mm ; pronotal medial length 0.50 mm ; pronotal width 0.87 mm ; metatibial length 0.58 mm ; length of forewing with scale 6.72 mm ; forewing width 2.24 mm . Integument dark brown, microsculpturing of head finely imbricate. Setation of head and body sparse; setae not evident on pronotum. Rs with four simple branches, Rs terminating before wing apex; M with four main branches, M terminating at wing apex, basalmost branch terminating at apical quarter of posterior wing margin; CuA pectinately branched, with at least four simple branches (remainder of CuA obscured in holotype ), cubital field occupying majority of posterior wing margin.