Afrotropical Asilidae (Diptera) 32. A Revision Of Anypodetus Hermann, 1907 With The Description Of Three New Species (Laphriinae)
Author
Londt, J. G. H.
Natal Museum, P. Bag 9070, Pietermaritzburg, South Africa
jlondt@nmsa.org.za
text
Annals of the Natal Museum
2000
2000-12-31
41
123
128
http://dx.doi.org/10.5281/zenodo.523759
journal article
10.5281/zenodo.523759
e927320e-7916-46b3-8bbb-dfc5b635c262
523759
2a
Wings banded with yellow -brown stained areas, membrane extensively covered with microtrichia; scutellar margin lacking setae; setae of prothoracic coxae and macrosetae of terga black; major mystacal macrosetae largely confined to epistomal margin. Illustrations - Head (
Fig. 1
), Antenna (
Fig. 4
), Wing (
Fig. 7
), Hypopygium (
Figs 16-18
), Ovipositor (
Fig. 34
), T2 (
Fig. 39
), Claws (
Fig. 45
), Distribution (
Fig. 48
) .....................................................................................
fascipennis
Engel
Synonymy
:
Anypodetus fascipennis
Engel, 1924:103
;
Anypodetus semirufus
Engel, 1924: 104
;
Anypodetus maculipennis
Ricardo, 1925: 242
.
Material examined
:
BOTSWANA
:
3♂
,
Farmer’s Brigade
,
Serowe
,
22
°
25'S
:
26
°
44'E
,
1000 m
,
28.xi.1980
,
P Forchhammer
,
Malaise trap
;
4♂
15♀
,
Farmer’s Brigade
,
Serowe
, 2226BD,
x.1985
(1♂2♀) -
xii.1985
(1♀) -
8.xi.1988
(1♀) -
9.xi.1988
(1♂1♀) -
x.1989
(5♀) -
xi.1989
(2♂5♀),
Forchhammer
,
Malaise trap
;
1♂
,
Farmer’s Brigade
,
6 km
SE
Serowe
,
1300 m
,
26.x.1984
,
Forchhammer
,
Forestry nursery
,
A. tortillis
woodland;
3♂
1♀
,
Serowe
,
26.xi.1982
(♂&♀)
21.xii.1982
(♂)
23.x.1982
(♂),
Forchhammer
, day;
1♀
,
Palapye
[
22
°
33'S
:
27
°
08'E
], B.P.,
18.x.1923
,
R Stevenson
(
SAMC
);
NAMIBIA
:
1♀
, SWA [=
South West Africa
= Namibia],
iii.1923
, Mus. Exp.;
1♀
,
Epupa Falls
[
16
°
55'S
:
13
°
10'E
],
Kunene
,
20-22.ii.1994
,
F Koch
(
ZMHB
);
1♀
,
Epupa Falls
17
°
00'S:13
°
15'E [use 16
°
55'S:13:10'E],
Kunene
,
19-21.ii.1994
,
H Schumann
(
ZMHB
);
3♀
,
Ruacana Falls
[
17
°
23'S
:
14
°
15'E
],
Kunene
,
23-24.ii.1994
,
F Koch
(
ZMHB
)
;
1♀
,
Kaross
[
19
°
30'S
:
14
°
20'E
],
ii.1925
, Mus. Exped. (
SAMC
);
1♂
,
27 mi
S
Tsumeb
[
19
°
14'S
:
17
°
43'E
],
10.xii.1956
;
1♀
,
Outjo
[
20
°
07'S
:
16
°
09'E
],
i.1925
,
Mus. Exped.
(
SAMC
);
1♂
1♀
,
80 mi
S
Gobabis
[
22
°
27'S
:
18
°
58'E
],
30.xii.1960
,
WD Haacke
;
1♀
,
Portsmut
,
Hakos Mts.
, 2416AB,
7.ii.1969
,
B Lamoral
;
1♂
,
Warmbad
[
28
°
27'S
:
18
°
44'E
],
Koakoveld
,
ii.1925
,
Mus. Exped.
(
SAMC
);
SOUTH AFRICA
:
1♂
1♀
,
Beacon Ranch
,
20 km
NW
Gravelotte
[
23
°
57'S
:
30
°
37'E
],
17.xi.1978
,
Brothers
&
J -Guillarmod
;
2♂
,
Moorddrift
[
24
°
17'S
:
28
°
57'E
],
7-19.x.1907
,
CJ Swierstra
;
1♂
1♀
,
Pretoria
[District
25
°
30'S
:
28
°
10'E
],
i.1919
,
Dr Brauns
;
1♂
,
Kaapmuiden
[
25
°
32'S
:
31
°
19'E
],
30.x.1918
,
RWE Tucker
(
SAMC
)
;
1♀
,
Vryburg
[
26
°
57'S
:
24
°
44'E
],
Griqualand West
,
x.1939
,
Mus. Staff
(
SAMC
)
;
1♂
1♀
,
De la Rey
[=
Delareyville
26
°
42'S
:
25
°
28'E
],
i.1917
(
♂
) -
15.i.1917
(
♀
),
Dr Brauns
;
11♂
4♀
,
De la Rey
,
i.1919
,
Dr Brauns
;
1♂
1♀
,
35 km
WNW
Upington
, 2820BD,
20.iii.1980
,
J Londt
&
L Schoeman
,
roadside vegetation
;
1♂
1♀
,
10 miles
W
Hopetown C.P.
[
29
°
37'S
:
24
°
05'E
],
27.i.1930
,
HK Munro
[
SANC
];
1♀
,
Bloemfontein
[District
29
°
10'S
:
26
°
00'E
],
12.xii.1920
,
HK Munro
;
1♂
,
22 km
SE
Graaff -Reinet on Pearston Rd.
,
32
°
27'S
:
24
°
38'E
,
750 m
,
7.xii.1989
,
J&H Londt
,
Open Karoo scrub/flowers
;
1♀
,
Merweville
[
32
°
40'S
:
21
°
31'E
] Dist.,
ii.1941
,
H. Zinn
(
SAMC
);
4♂
6♀
,
Willowmore
[
33
°
17'S
:
23
°
30'E
],
ii.1907
(♀) -
20.i.1908
(♀) -
1.i.1909
(♂) -
10.ii.1909
(♀) -
xii.1912
(♂) -
1.i.1913
(2♀) -
xii.1920
(♀) -
5.i.1922
(♂) -
10.xii.1922
(♂),
Dr Brauns
;
1♂
,
Willowmore
,
2.i.1913
,
Dr Brauns
(
SAMC
);
ZIMBABWE
:
1♂
2♀
,
Hillside
[
17
°
50'S
:
31
°
05'E
],
xi.1922
(♂) -
5.xi.1922
(♀) -
4.ii.1923
(♀),
Swinburne
&
Stevenson
;
1♂
,
Sawmills
[
19
°
35'S
:
28
°
02'E
],
14.xi.1924
,
RHR Stevenson
;
1♂
1♀
,
Bulawayo
[
20
°
09'S
:
28
°
35'E
],
xi.1922
(♂) - 1924 (♀),
RHR Stevenson
;
2♂
1♀
,
Bulawayo
,
12.x.
&
4.xi.1923
,
R. Stevenson
(
SAMC
).
Figs 10-21.
Anypodetus
species
, hypopygia. 10-12.
A. arachnoides
Oldroyd
(6 km N Vivo). 13-15.
A. fasciatus
Hermann (Serowe)
. 16-18.
A. fascipennis
Engel (Willowmore)
. 19-21.
A. leucothrix
sp. n.
(paratype). Scale lines = 1 mm.
Figs 22-33.
Anypodetus
species
, hypopygia. 22-24.
A. macroceros
sp. n.
(holotype). 25-27.
A. nigrifacies
Ricardo (Mkuzi)
.28-30.
A. phalaros
sp.n.
(37 kmNLouisTrichardt).31-33.
A. unicolor
Oldroyd
(20 mi S Ndumu). Scale lines = 1 mm.
Notes
: Type material not seen:
A. fascipennis
- 1♂, South Africa, Willowmore, 1.1.1907, Dr Brauns & 2♂1♀, Delarey, Dr Brauns, types in ZSMC;
A. semirufus
- 3♂1♀, South Africa, Willowmore, 1.I. - 5.I.1922, Dr Brauns.
Engel (1924)
did not indicate the type status of each specimen. If these are syntypes it will be necessary to designate lectotypes for both
fascipennis
and
semirufus
. I have seen many specimens from both Willowmore and Deleray collected by Brauns.
A. maculipennis
- 1♂1♀
syntypes
(
maculipennis
), Zimbabwe, Saw Mills (and other material from same locality) apparently in the National Museum of Zimbabwe, Bulawayo. I have seen other material from this locality.
Figs 34-36.
Anypodetus
species
, ovipositor, ventral aspect. 34.
A. fascipennis
Engel (Willowmore)
. 35.
A. nigrifacies
Ricardo (Mkuzi)
. 36.
A. unicolor
Oldroyd
(20 mi S Ndumu). Scale line = 1 mm.
Figs 37-44.
Anypodetus
species
, second abdominal tergum, distribution of pruinescence(unhatched).37.
A. arachnoides
Oldroyd.
38.
A. fasciatus
Hermann.
39.
A. fascipennis
Engel.
40.
A. leucothrix
sp. n.
41.
A. macroceros
sp.n.
42.
A. nigrifacies
Ricardo.
43.
A. phalaros
sp.n.
44.
A. unicolor
Oldroyd. Scale
line = 1 mm.
Figs 45-47.
Anypodetus
species.
45.
A. fascipennis
Engel
, tip of left hind tarsus. 46.
A. unicolor
Oldroyd
, egg dissected from female abdomen. 47.
A. leucothrix
sp. n.
, scutellum, dorsal view. Scale lines = 1 mm.
A. fascipennis
: Examination of the full series of specimens at my disposal supports acceptance of the synonymy of
semirufus
Engel
and
maculipennis
Ricardo
as suggested by
Oldroyd (1974)
. Although I have not seen Ricardo’s syntypes of
maculipennis
, I here designate the male as
lectotype
and the female as
paralectotype
.
Figs 48-49.
Anypodetus
species
distribution. 48.
A. fascipennis
Hermann
(closed circles),
A. leucothrix
sp. n.
(closed triangles),
A. nigrifacies
Ricardo
(closed squares),
A. phalaros
sp. n.
(open circles). 49.
A. arachnoides
Oldroyd
(closed triangles),
A. fasciatus
Hermann
(closed circles),
A.macroceros
sp. n.
(open circle),
A. unicolor
Oldroyd
(closed squares).
Fig. 50.
Anypodetus
, distribution of all data for genus, plotted on a map showing climatic regions. Abbreviations: D - Desert, M - Mediterranean, PS - Plateau slopes, SAP - Semi -arid plateau, SC - Subtropical coast, SL - Subtropical lowveld, SP - Subtropical plateau, TC - Temperate coast, TEP - Temperate eastern plateau, WTP - Warm temperate plateau.
TABLE 2
Distribution of
Anypodetus
species
in terms of climatic regions.? = doubtful record.
| Climatic Region (see below) |
| Species |
SP |
D |
M |
SAP |
TC |
TEP |
PS |
SC |
WTP |
SL
|
|
A. arachnoides
A. fasciatus
A. fascipennis
A. leucothrix
A. macroceros
A. nigrifacies
A. phalaros
A. unicolor
|
• • • - • - - - |
- - • • - - • - |
- - - - - - - - |
• • • • - - - - |
- - - - - - - - |
- • • - - - - •? |
- - • - - - • - |
- - - - - • - - |
• - • - - - - - |
- - - - - • - • |
| Records |
4 |
3 |
0 |
4 |
0 |
2?3 |
2 |
1 |
2 |
2 |
Abbreviations
:
SP - Subtropical plateau. TEP - Temperate eastern plateau.
D - Desert. PS - Plateau slopes.
M - Mediterranean. SC - Subtropical coast.
SAP - Semi -arid plateau. WTP - Warm temperate plateau.
TC - Temperate coast. SL - Subtropical lowveld.
A. fascipennis
: The most widely distributed species, known from six climatic regions. Not reported from the higher rainfall areas of the eastern parts of the subcontinent.