<document id="1AA8E8A654DF221681568D595C179535" ID="10.11646/zootaxa.4216.3.6" ID-CLB-Dataset="37356" ID-DOI="10.5281/zenodo.231847" ID-GBIF-Dataset="aaa31ce3-6f28-40b4-9ec8-0701a1e9791e" ID-ISSN="1175-5326" ID-Zenodo-Dep="231847" ID-ZooBank="60E45FC3-3B3B-44F0-B684-DB1F88F61CED" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1483689124128" checkinUser="plazi" docAuthor="Strümpher, Werner P. &amp; Scholtz, Clarke H." docDate="2017" docId="03AC5257FF8BB03640D92535FED0FEBD" docLanguage="en" docName="zootaxa.4216.3.6.pdf" docOrigin="Zootaxa 4216 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phoberus nyassicus Haaf, new status" docType="treatment" docVersion="8" lastPageNumber="297" masterDocId="FF952A2FFF8FB033404E2251FFB5FFC5" masterDocTitle="A new brachypterous species of Phoberus MacLeay (Coleoptera: Trogidae) and status changes of species in the East African P. nigrociliatus group" masterLastPageNumber="298" masterPageNumber="292" pageNumber="296" updateTime="1698435401945" updateUser="ExternalLinkService">
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<mods:title id="5F50442703D7D1BB60C61AAC16EB7493">A new brachypterous species of Phoberus MacLeay (Coleoptera: Trogidae) and status changes of species in the East African P. nigrociliatus group</mods:title>
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<mods:namePart id="E74F991055F36D495211267B107DDF73">Strümpher, Werner P.</mods:namePart>
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<mods:namePart id="A9F19676F7BC6BC245B7143ADC12F3C4">Scholtz, Clarke H.</mods:namePart>
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<mods:date id="39E4FA4C6F0CC087FCF8FAD9B9E8EB5B">2017</mods:date>
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<mods:identifier id="0248078B3045EF16C198EA38B31C5075" type="DOI">10.5281/zenodo.231847</mods:identifier>
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<treatment id="03AC5257FF8BB03640D92535FED0FEBD" ID-DOI="http://doi.org/10.5281/zenodo.6017606" ID-GBIF-Taxon="126094599" ID-Zenodo-Dep="6017606" LSID="urn:lsid:plazi:treatment:03AC5257FF8BB03640D92535FED0FEBD" httpUri="http://treatment.plazi.org/id/03AC5257FF8BB03640D92535FED0FEBD" lastPageId="5" lastPageNumber="297" pageId="4" pageNumber="296">
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<paragraph id="8BBAE341FF8BB03740D92535FDDDF8BB" blockId="4.[151,616,1892,1952]" box="[151,616,1892,1918]" pageId="4" pageNumber="296">
<heading id="D0F2542DFF8BB03740D92535FDDDF8BB" bold="true" box="[151,616,1892,1918]" fontSize="11" level="1" pageId="4" pageNumber="296" reason="1">
<emphasis id="B9713F53FF8BB03740D92535FDDDF8BB" bold="true" box="[151,616,1892,1918]" pageId="4" pageNumber="296">
<taxonomicName id="4C0598C2FF8BB03740D92535FE6BF8BB" ID-CoL="4GJX2" authority="Haaf" authorityName="Haaf" box="[151,478,1892,1918]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="296" phylum="Arthropoda" rank="species" species="nyassicus" status="stat. nov.">
<emphasis id="B9713F53FF8BB03740D92535FE33F8BB" bold="true" box="[151,390,1892,1918]" italics="true" pageId="4" pageNumber="296">Phoberus nyassicus</emphasis>
(Haaf)
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<taxonomicNameLabel id="A2428228FF8BB03741AB2535FDDDF8BB" box="[485,616,1892,1918]" pageId="4" pageNumber="296">new status</taxonomicNameLabel>
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</heading>
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<paragraph id="8BBAE341FF8BB03740D925D6FE94F865" blockId="4.[151,616,1892,1952]" box="[151,289,1927,1952]" pageId="4" pageNumber="296">
(
<figureCitation id="133EFFC4FF8BB03740D125D6FF40F865" box="[159,245,1927,1952]" captionStart="FIGURE 5" captionStartId="5.[151,250,1121,1143]" captionTargetBox="[308,1307,451,1080]" captionTargetId="figure@5.[279,1307,431,1099]" captionTargetPageId="5" captionText="FIGURE 5. A, Dorsal habitus of Phoberus nyassicus; B, dorsal and lateral views of the aedeagus of P. nyassicus; C, elytral sculpture of P. nyassicus; D, metathoracic wing of P. nyassicus." httpUri="https://zenodo.org/record/231852/files/figure.png" pageId="4" pageNumber="296">Fig. 5A</figureCitation>
–D)
</paragraph>
</subSubSection>
<subSubSection id="C31FB0CAFF8BB03640D92581FD42FF2E" lastPageId="5" lastPageNumber="297" pageId="4" pageNumber="296" type="reference_group">
<paragraph id="8BBAE341FF8BB03640D92581FD42FF2E" blockId="4.[151,604,1999,2021]" lastBlockId="5.[151,920,152,235]" lastPageId="5" lastPageNumber="297" pageId="4" pageNumber="296">
<treatmentCitationGroup id="AB15C46FFF8BB03640D92581FD42FF2E" lastPageId="5" lastPageNumber="297" pageId="4" pageNumber="296">
<taxonomicName id="4C0598C2FF8BB03740D92581FDE9F820" ID-CoL="59DNM" authority="Haaf, 1953: 330" authorityName="Haaf" authorityPageNumber="330" authorityYear="1953" box="[151,604,1999,2021]" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="296" phylum="Arthropoda" rank="subSpecies" species="nigrociliatus" subSpecies="nyassicus">
<emphasis id="B9713F53FF8BB03740D92581FE07F820" box="[151,434,1999,2021]" italics="true" pageId="4" pageNumber="296">Trox nigrociliatus nyassicus</emphasis>
<treatmentCitation id="0AA4C550FF8BB03741F7259EFDE9F820" author="Haaf" box="[441,604,1999,2021]" page="330" pageId="4" pageNumber="296" year="1953">
<bibRefCitation id="EF949EB0FF8BB03741F7259EFD98F820" author="Haaf" box="[441,557,1999,2021]" pageId="4" pageNumber="296" refString="Haaf, E. (1953) Die afrikanischen Arten der Gattung Trox F. (Col. Scarab.). Entomologische Arbeiten aus dem Museum Georg Frey, 4, 309 - 346." type="journal article" year="1953">Haaf, 1953</bibRefCitation>
: 330
</treatmentCitation>
</taxonomicName>
<taxonomicName id="4C0598C2FF8AB03640D922C8FD06FF6B" ID-CoL="59DNM" authority="Scholtz 1980: 93" authorityName="Scholtz" authorityPageNumber="93" authorityYear="1980" box="[151,691,152,174]" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="297" phylum="Arthropoda" rank="subSpecies" species="nigrociliatus" subGenus="Trox" subSpecies="nyassicus">
<emphasis id="B9713F53FF8AB03640D922C8FF71FF6B" box="[151,196,153,174]" italics="true" pageId="5" pageNumber="297">Trox</emphasis>
(
<emphasis id="B9713F53FF8AB036409D22C8FF4AFF6B" box="[211,255,153,174]" italics="true" pageId="5" pageNumber="297">Trox</emphasis>
) 
<emphasis id="B9713F53FF8AB036414122C9FE42FF6B" box="[271,503,152,174]" italics="true" pageId="5" pageNumber="297">nigrociliatus nyassicus</emphasis>
: 
<treatmentCitation id="0AA4C550FF8AB036424A22C9FD06FF6B" author="Scholtz" box="[516,691,152,174]" page="93" pageId="5" pageNumber="297" year="1980">
<bibRefCitation id="EF949EB0FF8AB036424A22C9FD38FF6B" author="Scholtz" box="[516,653,152,174]" pageId="5" pageNumber="297" refString="Scholtz, C. H. (1980) Monograph of the genus Trox F. (Coleoptera: Trogidae) of Subsaharan Africa. Cimbebasia Memoir, 4, 1 - 104." type="journal article" year="1980">Scholtz 1980</bibRefCitation>
: 93
</treatmentCitation>
</taxonomicName>
; 
<treatmentCitation id="0AA4C550FF8AB03642F322C9FCE8FF6B" author="Scholtz" box="[701,861,152,174]" page="5" pageId="5" pageNumber="297" year="1982">
<bibRefCitation id="EF949EB0FF8AB03642F322C9FCF3FF6B" author="Scholtz" box="[701,838,152,174]" pageId="5" pageNumber="297" refString="Scholtz, C. H. (1982) Catalogue of world Trogidae (Coleoptera: Scarabaeoidea). Entomology Memoir, Department of Agriculture and Fisheries, Republic of South Africa, 54, 1 - 27." type="journal article" year="1982">Scholtz 1982</bibRefCitation>
: 5
</treatmentCitation>
<taxonomicName id="4C0598C2FF8AB03640D922E9FD46FF09" ID-CoL="59DNM" authority="Scholtz 1986: 361" authorityName="Scholtz" authorityPageNumber="361" authorityYear="1986" box="[151,755,182,205]" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="297" phylum="Arthropoda" rank="subSpecies" species="nigrociliatus" subGenus="Phoberus" subSpecies="nyassicus">
<emphasis id="B9713F53FF8AB03640D922E9FF71FF08" box="[151,196,184,205]" italics="true" pageId="5" pageNumber="297">Trox</emphasis>
(
<emphasis id="B9713F53FF8AB036409D22E7FE81FF09" box="[211,308,182,204]" italics="true" pageId="5" pageNumber="297">Phoberus</emphasis>
) 
<emphasis id="B9713F53FF8AB036410D22E7FD9EFF08" box="[323,555,182,205]" italics="true" pageId="5" pageNumber="297">nigrociliatus nyassicus</emphasis>
: 
<treatmentCitation id="0AA4C550FF8AB036427622E7FD46FF09" author="Scholtz" box="[568,755,182,204]" page="361" pageId="5" pageNumber="297" year="1986">
<bibRefCitation id="EF949EB0FF8AB036427622E7FD75FF09" author="Scholtz" box="[568,704,182,204]" pageId="5" pageNumber="297" refString="Scholtz, C. H. (1986) Phylogeny and systematics of the Trogidae (Coleoptera: Scarabaeoidea). Systematic Entomology, 11, 355 - 363." type="journal article" year="1986">Scholtz 1986</bibRefCitation>
: 361
</treatmentCitation>
</taxonomicName>
; 
<treatmentCitation id="0AA4C550FF8AB03642B322E7FC2DFF09" author="Zidek" box="[765,920,182,204]" page="13" pageId="5" pageNumber="297" year="2013">
<bibRefCitation id="EF949EB0FF8AB03642B322E7FCC0FF09" author="Zidek" box="[765,885,182,204]" pageId="5" pageNumber="297" refString="Zidek, J. (2013) Checklist and bibliography of the Trogidae (Coleoptera: Scarabaeoidea). Insecta Mundi, 314, 1 - 38." type="journal article" year="2013">Zidek 2013</bibRefCitation>
: 13
</treatmentCitation>
<taxonomicName id="4C0598C2FF8AB03640D92284FD42FF2E" ID-CoL="4GJX2" authority="Strumpher et al. 2016: 78" authorityName="Strumpher et al." authorityPageNumber="78" authorityYear="2016" box="[151,759,213,235]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="297" phylum="Arthropoda" rank="subSpecies" species="nigrociliatus" subSpecies="nyassicus">
<emphasis id="B9713F53FF8AB03640D92284FE53FF2E" box="[151,486,213,235]" italics="true" pageId="5" pageNumber="297">Phoberus nigrociliatus nyassicus</emphasis>
: 
<treatmentCitation id="0AA4C550FF8AB03641BD2284FD42FF2E" author="Strumpher" box="[499,759,213,235]" page="78" pageId="5" pageNumber="297" year="2016">
<bibRefCitation id="EF949EB0FF8AB03641BD2284FD66FF2E" author="Strumpher" box="[499,723,213,235]" pageId="5" pageNumber="297" refString="Strumpher, W. P., Villet, M. H., Sole, C. L. &amp; Scholtz, C. H. (2016) Overview and revision of the extant genera and subgenera of Trogidae (Coleoptera: Scarabaeoidea). Insect Systematics and Evolution, 47, 53 - 82. https: // doi. org / 10.1163 / 1876312 X- 46052133" type="journal article" year="2016">
Strümpher 
<emphasis id="B9713F53FF8AB036422A2287FD25FF2E" box="[612,656,213,235]" italics="true" pageId="5" pageNumber="297">et al</emphasis>
. 2016
</bibRefCitation>
: 78
</treatmentCitation>
</taxonomicName>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C31FB0CAFF8AB03640D92346FED0FEBD" pageId="5" pageNumber="297" type="diagnosis">
<paragraph id="8BBAE341FF8AB03640D92346FED0FEBD" blockId="5.[151,1436,279,376]" pageId="5" pageNumber="297">
<emphasis id="B9713F53FF8AB03640D92346FEA4FEF5" bold="true" box="[151,273,279,304]" pageId="5" pageNumber="297">Diagnosis.</emphasis>
<taxonomicName id="4C0598C2FF8AB03641592346FE40FEF5" box="[279,501,279,304]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="297" phylum="Arthropoda" rank="species" species="nyassicus">
<emphasis id="B9713F53FF8AB03641592346FE40FEF5" box="[279,501,279,304]" italics="true" pageId="5" pageNumber="297">Phoberus nyassicus</emphasis>
</taxonomicName>
is very similar to 
<taxonomicName id="4C0598C2FF8AB03642F22348FC8EFEF5" box="[700,827,279,304]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="297" phylum="Arthropoda" rank="species" species="nyikanus">
<emphasis id="B9713F53FF8AB03642F22348FC8EFEF5" box="[700,827,279,304]" italics="true" pageId="5" pageNumber="297">P. nyikanus</emphasis>
</taxonomicName>
in appearance, but can be distinguished from the latter by distinct, evenly spaced tubercles on the elytra; 
<taxonomicName id="4C0598C2FF8AB03642A8236CFCDCFE90" box="[742,873,316,341]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="297" phylum="Arthropoda" rank="species" species="nyikanus">
<emphasis id="B9713F53FF8AB03642A8236CFCDCFE90" box="[742,873,316,341]" italics="true" pageId="5" pageNumber="297">P. nyikanus</emphasis>
</taxonomicName>
has the tubercles partially fused, forming semicontinuous ridges.
</paragraph>
</subSubSection>
</treatment>
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