A key to the Russian species of the genus Coenosia Meigen (Diptera, Muscidae), with the description of one new species and new synonymies Author Sorokina, Vera S. text Zootaxa 2023 2023-12-18 5389 1 79 107 https://www.mapress.com/zt/article/download/zootaxa.5389.1.4/52487 journal article 10.11646/zootaxa.5389.1.4 1175-5326 10404495 9BD1B8FF-28B7-4D6E-8F4A-FF1E79C30DDB Key to males 1. Costa extending only to, or very slightly beyond, apex of R 4+5 .................................................. 2 - Costa extending to apex of M 1 ........................................................................... 3 2. Lower calypter not longer than upper one. Postpedicel with a spine-like dorsal tip.—In Asia only Siberia and China ; Eurasian................................................................................... connectens (Hennig) - Lower calypter much longer than upper one. Postpedicel without a spine-like dorsal tip, with a more or less rounded tip.— European Russia , Siberia, Far East; Palaearctic.............................................. agromyzina (Fallén) 3. Hind tibia with a median anterior seta and an anterodorsal seta at almost the same level and close together ( Fig. 9A )....... 4 - Hind tibia without an anterior seta at middle close to the anterodorsal seta, but an anteroventral seta present............. 9 4. Scutum, scutellum and abdomen shining black.—European Russia ; Palaearctic.......................... atra (Meigen) - Scutum, scutellum and abdomen not shining black........................................................... 5 5. Parafacial at least as broad as postpedicel; longest aristal hairs at least as long as width of postpedicel; outer vertical setae longer and stronger than postocular setulae.—European Russia , Siberia; Holarctic......................... tigrina (Fabricius) - Parafacial narrower than width of postpedicel; longest aristal hairs much shorter than width of postpedicel; outer vertical setae at most as long as postocular setulae...................................................................... 6 6. All femora predominantly black.—European Russia ; Holarctic, Oriental.............................. humilis Meigen - Femora yellow, at most fore femur dorsally and mid femur apically brownish...................................... 7 7. Antenna entirely dark or flagellomere partly yellow; frontal vitta less dusted, more contrasting with fronto-orbital plates; abdomen with a midline and paired brown spots.—European Russia , Siberia, Far East; Palaearctic, Afrotropical, Oriental........................................................................................... strigipes Stein - Antenna entirely yellow; frontal vitta densely silvery white to yellow dusted, hardly differentiated from fronto-orbital plates; abdomen grey dusted, at most with indistinct pattern.—European Russia ; Holarctic, Neotropical, Afrotropical, Oriental, Australasian............................................................................. attenuata Stein 9. Last tarsomere at least of fore legs enlarged and black, contrasting with rest of tarsi which are yellow ( Fig. 9C ).......... 10 - Last tarsomere not dilated on any pair of legs.............................................................. 11 10. Last tarsomere on all legs dilated and black, contrasting with rest of yellow tarsi; one proepisternal seta; abdomen ovate.—Only European Russia ......................................................................... rhaensis Hennig - Only last tarsomere of fore legs enlarged and black, contrasting with rest of yellow tarsi; two proepisternal seta; abdomen unusually long and narrow with nearly parallel sides ( Fig. 8A ). Terminalia in Figs 3 A–C .— Georgia , Russian Caucasus............................................................................................ ozerovi Vikhrev 11. Hind tibia with strong and long preapical dorsal and posterodorsal setae close to each other ( Fig. 9B ).................. 12 - Hind tibia without strong and long close approximated preapical dorsal and posterodorsal setae...................... 13 12. Mid and hind femora and all tibia predominantly yellow.—European Russia , Siberia, Far East; Eurasian..................................................................................................... intermedia (Fallén) - Mid and hind femora and all tibia predominantly black but fore and mid trochanters yellow.—European Russia ; European........................................................................................... means Meigen 13. Lower calypter much shorter or scarcely longer than upper one................................................ 14 - Lower calypter much longer than upper one............................................................... 38 14. Oral vibrissae yellow or golden; head almost square in lateral view; frons reddish above antenna, postpedicel black, very long, 4 times as long as wide ( Fig. 10E ).— Norway , NE Russia , Far East; Holarctic........................ flaviseta (Huckett) - With a different combination of characters................................................................ 15 15. Basal scutellar setae reduced, apical setae weak and short.................................................... 16 - Both pairs of scutellars strong and long................................................................... 17 16. Body predominantly bluish-gray ( Fig. 7A ); mid femur with anteroventral setae.—East Siberia and Far East; Holarctic.............................................................................................. ciliata Hennig - Body predominantly yellow; mid femur without anteroventral setae.—Only European Russia ........... flavissima Hennig 17. All femora black, at most with apices narrowly yellow....................................................... 18 - At least mid and hind femora yellow on basal 1/3........................................................... 23 18. Antenna including both basal segments yellow; hind tibia without a submedian posterodorsal seta.—European Russia , East Siberia; Eurasian..................................................................... dealbata (Zetterstedt) - Antenna entirely black; hind tibia with or without a distinct submedian posterodorsal seta........................... 19 19. Ventral surface of mid and hind femora and tibia with numerous long semi-erect hairs; posterior seta on fore tibia and anterodorsal setae on mid and hind tibiae very long, at least as long as half of length of tibia ( Fig. 7C ). Terminalia in Figs 6 A–E .—European Russia ,?Far East; Oriental and Palaearctic ( Japan ).............................. penicillata Hennig - Ventral surface of mid and hind legs without numerous long semi-erect hairs; setae on tibiae not longer than one-third third of tibial length......................................................................................... 20 20. Smaller species ( 3.5–4.7 mm ); all tibiae predominantly black, only narrowly yellow basally; hind femur with 2–3 posteroventral setae on basal half; hind tibia without or with a very short posterodorsal, about 1/3 the length of the anterodorsal seta..... 21 - Larger species (5.0– 6.5mm ); all tibiae brownish-yellow or yellow; hind femur with a full row of strong and long posteroventrals; posterodorsal seta of hind tibia long and only a little shorter than anterodorsal.................................... 22 21. Mid femur with a preapical anterior seta; mid tibia with a long strong anterodorsal seta, at least as long as posterodorsal seta; abdomen cylindrical, with weak brownish rounded spots; scutum with two distinct brown vittae ( Fig. 7D ). Terminalia in Figs 3 D, E .—Only Siberia and China ; Asian................................................. subgracilis Xue & Cui - Mid femur without a preapical anterior seta; mid tibia with a weak anterodorsal seta which is much shorter than posterodorsal seta; abdomen oval, with pairs of indistinct brown rectangular spots and brownish dust between them; scutum with 3 distinct brown longitudinal vittae and brownish dust between dorsocentral setae (Figures in Sorokina 2022 ).—Only North of Russian Far East................................................................................. galina Sorokina 22. Cerci narrow and long ( Figs 6 F, G ). All tibiae yellow ( Fig. 7F ); abdomen with rounded spots.—Only European Russia , Siberia and Russian Far East....................................................................... oralis Schnabl - Cerci short and wide ( Figs 5 E, F ). All tibiae brownish-yellow, darkened in apical 2/3–4/5, sometimes completely yellow ( Fig. 7H ); abdomen with more or less triangular spots.—Only West Siberia and Russian Far East..... ukokensis Sorokina (part) 23. Lower calypter extremely small, shorter than the upper one, strip-like........................................... 24 - Lower calypter not strip-like, slightly protruding beyond margin of upper one.................................... 25 24. Smaller size: 3.5 mm ( Fig. 10D ). Mid femur without a strong anterior preapical seta; palpus yellow; mid and hind femora fuscous on distal half. Terminalia in Figs 6 H, I .—West Siberia, Far East; Holarctic.................... tendipes Huckett - Larger size: 5–6 mm ( Fig. 10A ). Mid femur with a strong anterior preapical seta; palpus completely dark or yellow with dark tip; mid and hind femora yellow, with a very small dark spot at apex.—Siberia, Far East; Holarctic.................................................................................................... baicalensis Hennig (part) 25. At least apical half or third of hind femur black or fuscous on dorsal part of apex.................................. 26 - All femora completely yellow.......................................................................... 29 26. Scutum with 3 distinct brown narrow longitudinal vittae along dorsocentral and acrostichal setae, with brown dust between dorsocentral setae, reaching the apex of scutellum; mid tibia with very short anterodorsal seta which shorter then posterodorsal seta; mid and hind femora yellow on basal ½-1/3, at least mid and hind tibiae darkened (Figures in Sorokina 2022 ).—Only North of Russian Far East............................................................. wrangelensis Sorokina - Scutum with 2 distinct brown narrow longitudinal vittae along dorsocentral setae, without brown dust; mid tibia with long and subequal anterodorsal and posterodorsal setae; mid and hind femora yellow at least on basal 2/3, all tibiae yellow........ 27 27. Fore femur darkened at most on dorsal surface, ventral surface yellow; mid and hind femora only with a small dark dorsal mark (Figures of C. morrisoni in Sorokina 2022 ).—West Siberia, Far East; Holarctic............ cingulipes (Zetterstedt) (part) - Fore femur completely black; mid and hind femur with black apical ring........................................ 28 28. Hind tibia with a well-developed submedian posterodorsal seta, as long as 2/3 of anterodorsal seta; cheek higher than width of postpedicel ( Fig. 9G ); sternite 5 small and short, as long as length of sternite 4; cerci short and wide. Larger size: 5.0– 6.8 mm .—Far East; Holarctic................................................................ alaskensis Huckett - Hind tibia without submedian posterodorsal seta, or with a short weak seta which shorter than anterodorsal seta; cheek not higher than width of postpedicel ( Fig. 4I ); sternite 5 large, as long as length of sternite 3+4; cerci long and thin. Smaller size: 3.0– 5.5 mm .—European Russia , Siberia, Far East; Holarctic.............................. octopunctata (Zetterstedt) 29. Mid femur in apical third without a strong anterior preapical seta.............................................. 30 - Mid femur in apical third with a strong anterior preapical seta................................................. 34 30. Body entirely yellow; scutum densely grey dusted, pleura yellow with a grey dusted spot on anepimeron; apical tarsomeres of all legs blackish, hind tibia with 4 long and thin posteroventrals in apical third.—Only Russian Far East (Kunashir Island).......................................................................................... polina Vikhrev - At least thorax and abdomen grey, not yellow.............................................................. 31 31. Antenna and palpus partly or completely yellow; scutum brownish dusted, without dark longitudinal stripes (Figures in Vikhrev & Sorokina 2018 ).—West Siberia, Far East; Holarctic.......................................... demoralis Huckett - Antenna and palpus wholly dark; scutum with brownish longitudinal stripes...................................... 32 32. Scutum with thick brown dust between dc which reaches the apex of scutellum; abdomen with brown rectangular marks on tergites 1+2–5 which merge in two stripes; mid femur with apical posteroventral seta; two proepisternals (Figures in Sorokina 2022 ).—Only North of Russian Far East................................................... koni Sorokina (part) - Scutum with 2–3 more or less distinct longitudinal stripes, without brown dust between dc ; abdomen with small rounded spots on tergites 1+2–5, sometimes indistinct; mid femur without apical posteroventral seta; usually one proepisternal......... 33 33. Scutum with two distinct brown longitudinal stripes; height of gena 1.3–1.5 times width of postpedicel; mid tibia with short anterodorsal seta, which is half as long as posterodorsal seta.—European Russia , Siberia, Far East; Eurasian.......................................................................................... bilineella (Zetterstedt) (part) - Scutum with three indistinct brownish longitudinal stripes; height of gena 2.0–2.5 times width of postpedicel ( Fig. 10G ); mid tibia with long anterodorsal seta, which is 2 times as long as posterodorsal seta. Terminalia in Figs 3H, I .—Siberia, Far East; Holarctic......................................................................... conflicta Huckett (part) 34. Antenna and palpus wholly dark; mid tibia with 1 anterodorsal seta. Larger size: 5–6 mm ........................... 35 - Antenna and palpus partly or completely yellow; mid tibia without anterodorsal seta. Smaller size: 3–5 mm ............ 36 35. Abdomen slender, elongate, cylindrical, with a pair of narrow oblong marks on tergites 1+2–5; cheek always higher than width of postpedicel ( Fig. 10A ).—Siberia, Far East; Holarctic................................. baicalensis (Schnabl) (part) - Abdomen stout and conical, with a pair of triangular spots on tergites 1+2–5; cheek higher or not than width of postpedicel; mid and hind femur only with small dark dorsal mark (Figures to C. morrisoni in Sorokina 2022 ).—West Siberia, Far East; Holarctic.................................................................... cingulipes (Zetterstedt) (part) 36. Postpronotal lobe and greater part of pleura yellow, abdomen yellow with dark spots on tergites 3–5 (sometimes only on tergite 4). Terminalia in Figs 3 F, G .—Only South of Russian Far East................................... ghilarovi Lobanov - Postpronotal lobe and all pleura gray, abdomen yellow or mostly grey.......................................... 37 37. Palpus at least broadly yellow at tip, usually almost or completely yellow; tip of antennal pedicel yellow; dark dorsocentral vittae weakly visible; sternite 5 with setae and setulae long and dense.—European Russia , Siberia, Far East; Holarctic, Oriental........................................................................................ mollicula (Fallén) - Palpus wholly dark or at least dark at tip; antennal pedicel and flagellomere usually narrowly reddish basally; dark dorsocentral vittae conspicuous and strongly marked; sternite 5 with setae and setulae shorter and sparser.—Siberia, Far East; Eurasian....................................................................................... alpicola (Pokorny) 38. Postpedicel with a spine-like dorsal tip ( Fig. 9D )........................................................... 39 - Postpedicel without a spine-like dorsal tip, rounded at tip..................................................... 44 39. Sternite 5 with dense long yellow setae; antenna and vibrissa completely yellow ( Fig. 9E ).—Only Siberia and Mongolia ; Asian................................................................................. kosterini Vikhrev - Sternite 5 without dense long yellow setae; antenna and vibrissa dark........................................... 40 40. All femora completely reddish-yellow.—European Russia , Siberia; Eurasian........................ ambulans Meigen - Femora at least partly black............................................................................ 41 41. Legs black, at most knees pale.......................................................................... 42 - At least mid and hind femora pale basally................................................................. 43 42. Body uniformly whitish-blue dusted; frontal vitta thrice as wide as fronto-orbital plate; postpedicel not reaching to the level of oral margin; height of gena equal to width of postpedicel.—European Russia ; European........... flavimana (Zetterstedt) - Body whitish-grey, with a dark pattern; frontal vitta twice as wide as fronto-orbital plate; postpedicel long, reaching to the level of oral margin; height of gena 1.5–2.0 times width of postpedicel ( Fig. 1A ). Terminalia in Figs 1 D–F .—Only North of Russian Far East ( Magadan )......................................................................... luxia sp. nov. 43. Femora predominantly yellow: fore femur with dark dorsal streak, mid and hind femora with darkened apical third.—European Russia ; European........................................................................ acuminata Strobl - Fore femora predominantly dark, mid and hind femora usually darkened in distal halves or more.—European Russia , Siberia, Far East; Holarctic.................................................................. pulicaria (Zetterstedt) 44. Only one proepisternal seta (sometimes with an additional weak seta on one side)................................. 45 - Two proepisternals................................................................................... 51 45. Mid tibia with 1 strong anterodorsal seta.................................................................. 46 - Mid tibia without an anterodorsal seta.................................................................... 48 46. Scutum with two distinct brown longitudinal stripes; postpedicel ca. 4 times as long as wide; mid tibia of male with a short anterodorsal seta, which is half as long as posterodorsal seta; pulvilli elongate.—European Russia , Siberia, Far East; Eurasian............................................................................ bilineella (Zetterstedt) (part) - Scutum without or with three indistinct brownish longitudinal stripes; postpedicel ca. 2.0–3.0 times as long as wide; mid tibia of male with a long anterodorsal seta, which is 2 times as long as posterodorsal seta; pulvilli small.................... 47 47. Height of gena 2.0–2.5 times width of postpedicel; antenna black or postpedicel brown at base ( Fig. 10G ).—Siberia, Far East; Holarctic......................................................................... conflicta Huckett (part) - Height of gena less than 2 times width of postpedicel; postpedicel yellow on basal half, pedicel yellow at apex.—Far East; Holarctic......................................................................... incisurata van der Wulp 48. Frons wide, at middle 0.5 of head-width at this point; frontal vitta and fronto-orbital plate velvety black; scutum and scutellum densely brown dusted; postpedicel black, long, ca. 3.5 times as long as wide and almost reaching mouth-edge (Figures in Sorokina 2022 ). Smaller size: 2.1–2.2 mm .—Only Siberia and Russian Far East...................... adriani Sorokina - Frons narrower; frontal vitta and a fronto-orbital plate not velvety black, grey; scutum and scutellum without dense brown dust, grey with brownish stripes; postpedicel black or reddish at base, small, not reaching mouth-edge. Larger size: 2.5–5.0 mm... .................................................................................................. 49 49. Posterior and posteroventral setulae on fore tarsomeres elongate and semi-erect, longer than depth of tarsus; mid femur usually with 2 preapical posterodorsals and 3-4 strong posteroventrals in basal 2/3.—European Russia ; European.................................................................................................... nigridigita Rondani - Posterior and posteroventral setulae on fore tarsomeres short, decumbent; setae on mid femur different................ 50 50. Hind femur with one or more well developed posteroventral setae on apical half; postpedicel more or less yellowish.—European Russia , Siberia, Far East; Eurasian.......................................................... rufipalpis Meigen - Hind femur with distinct posteroventrals on basal half only; postpedicel black.—European Russia , Siberia; Palaearctic................................................................................. testacea (Robineau-Desvoidy) 51. All femora black, only narrowly yellow apically............................................................ 52 - Femora at least partly yellow........................................................................... 56 52. Tibiae light yellow to reddish-yellow; sternite 5 projecting ventrally from abdomen at an angle of about 45º; hind femur with 2–3 long anteroventral setae and with 1 long posteroventral seta at middle.—European Russia , Siberia; Palaearctic..................................................................................... Macrorchis meditata (Fallén) - Tibiae dark yellow to brownish or darkened to almost black, only yellow on basal third; sternite 5 normal, not projecting ventrally below abdomen; hind femur with more or less full rows of anteroventrals and posteroventrals................ 53 53. Hind tibia without submedian posterodorsal seta; only one pair of presutural dorsocentrals.......................... 54 - Hind tibia with submedian posterodorsal; two pair of presutural dorsocentrals.................................... 55 54. Abdomen short, ovate, enlarged dorso-ventrally ( Fig. 4A ); mid tibia with anterodorsal seta. Terminalia in Figs 5 A–C .— European Russia ; European............................................................ nevadensis Lyneborg - Abdomen slender and long, nearly parallel-sided ( Fig. 4G ); mid tibia usually without anterodorsal seta.—European Russia ; European............................................................................ octosignata Rondani 55. Scutum with 3 distinct brown longitudinal vittae; lower calypter much longer than upper one; usually calypters white ( Fig. 8D ). Terminalia in Fig. 5 D .—Only West Siberia and Russian Far East................................ nigrotincta Hennig - Scutum with 2 very diffuse brownish longitudinal vittae; lower calypter scarcely longer than upper one; usually calypters yellow ( Figs 7H , 8E ). Terminalia in Figs 5 E, F .—Only West Siberia and Russian Far East...... ukokensis Sorokina (part) 56. Fore femur predominantly dark......................................................................... 57 - Fore femur predominantly yellow or at most with a dark dorsal streak or spot..................................... 68 57. Two pairs of presutural dorsocentral setae, the anterior pair about half as long as posterior pair....................... 58 - Only one pair of presutural dorsocentrals................................................................. 62 58 Smaller size: 2.8–3.0 mm. Hind tibia black; hind tarsi thickened, all segments about twice as wide as those of mid tarsus; fore tarsomeres 1–4 yellow, tarsomere 5 black ( Fig. 11A ).—Far East; Holarctic..................... atritibia Ringdahl (part) - Larger size: 4.2–6.2. Hind tibia yellow; hind tarsomeres slender, mostly as wide as mid tarsomeres................... 59 59. Hind femur only in distal third with 2–3 long anteroventral setae, long posteroventral setae only in basal third; mid and hind femora predominantly yellow, only with a weak brownish subapical ring.—Only Siberia and Russian Far East.............................................................................................. sibirica Hennig (part) - Hind femur with full rows of long and strong anteroventrals and posteroventrals; mid and hind femora black at least in distal ½ or 1/3.............................................................................................. 60 60. Mid and hind femora mainly black, with yellow only basal 1/4 or 1/3; mid tibia without strong posterodorsal setae; abdomen short, as long as thorax; sternites II–IV strongly shortened, together shorter than sternite V. Body length 4.5 mm .—West Siberia; Holarctic............................................................................... comita (Huckett) - Mid and hind femora yellow at least in basal half; mid tibia with strong or without hair-like posterodorsal seta; abdomen oblong, longer than thorax; sternites II–IV not shortened, together longer than sternite V. Body length 5.0– 6.2 mm ....... 61 61. Hind tibia without strong or with hair-like posterodorsal setae in middle; mid and hind femora yellow in basal ½ or 2/3; antenna occupying 2/3 of height of face ( Fig. 11C ); epandrium without apical process. Terminalia in Fig. 5 G .—Only Siberia and Russian Far East....................................................................... apukaensis Hennig - Hind tibia with 1 or 2 strong posterodorsal setae in middle; mid and hind femora yellow in basal 2/3 or 3/4; antenna occupying 3/4 of height of face ( Fig. 11F ); epandrium with apical process. Terminalia in Fig. 5 H .—Only Russian Far East and China ; Asian.................................................................................... xuei Cui et Li 62. Sternite 5 very large, projecting ventrally from abdomen, with coarse recurved setae (Figures in Sorokina 2022 ).—Only West Siberia.............................................................................. tschernovi Sorokina - Sternite 5 small, not projecting ventrally from abdomen, without coarse setae..................................... 63 63. Sternite 5 with long and fine hairs; hind femur without anteroventral and posteroventral setae, but with long and fine hairs on their place.—European Russia ; European................................................ dubiosa Hennig (part) - Sternite 5 without long and fine hairs; at least 1 or 2 strong anteroventral and posteroventral setae and with only a few fine hairs near base on hind femur.............................................................................. 64 64. Hind tibia black; hind tarsomeres thickened, all segments about twice as wide as those of mid leg; fore tarsomeres 1–4 yellow, tarsomere 5 contrastingly black ( Fig. 11A ).—Far East; Holarctic............................. atritibia Ringdahl (part) - Hind tibia yellow or brown; hind tarsomeres slender, mostly as wide as those of mid leg; fore tarsus with at least tarsomeres 3–5 darkened....................................................................................... 65 65. Scutum with 3 distinct brown narrow longitudinal vittae along dorsocentral and acrostichal setae, with brown dust between dorsocentral setae, reaching the apex of scutellum; mid femur with apical posteroventral setae; gena higher than width of postpedicel, 2 times as long as postpedicel; mid and hind femora usually yellow on basal 1/3 (Figures in Sorokina 2022 ).—Only North of Russian Far East........................................................ wrangelensis Sorokina (part) - Scutum without distinct longitudinal vittae and brown dust between dorsocentral setae, scutellum grey; mid femur without apical posteroventral setae; gena not higher than width of postpedicel; mid and hind femora yellow at least on basal 2/3... 66 66. All tarsomeres black; mid femur in apical third with preapical anterior seta; cerci thin and long.—European Russia ; European.......................................................................... femoralis (Robineau-Desvoidy) - At least fore basitarsus yellowish; mid femur in apical third without preapical anterior seta; cerci not long, wide......... 67 67. Only last tarsomere of fore leg black; mid femur entirely yellow; hind femur darkened at apex.—European Russia , Siberia, Far East; Eurasian....................................................................... pygmaea (Zetterstedt) - Apical darkening of fore tarsus not confined to last tarsomere; mid femur with indistinct and hind femur with distinct subapical dark ring.—European Russia , Siberia, Far East; Holarctic.......................................... verralli Collin 68. Postpedicel yellow at least basally....................................................................... 69 - Antenna entirely black................................................................................ 71 69. Hind tibia with long and strong submedian posterodorsal seta.—European Russia , Siberia, Far East; Eurasian................................................................................................. flavicornis (Fallén) - Hind tibia without long and strong submedian posterodorsal seta............................................... 70 70. Mid tibia with anterodorsal seta; anteroventral seta on hind tibia in a normal, truly anteroventral position; abdomen at least basally more or less yellow; parafacial reaching about ½ width of postpedicel.—Siberia; Eurasian....................................................................................................... ruficornis (Macquart) - Mid tibia without anterodorsal seta; long and fine anteroventral seta on hind tibia in strictly ventral position; abdomen entirely dark; parafacial narrow, reaching about 1/3 width of postpedicel.—European Russia ; European........... pudorosa Collin 71. Apical scutellar setae distinctly shorter and weaker than basal scutellars......................................... 72 - Apical and basal scutellars subequal in length.............................................................. 73 72. Axillary lobe of wing less developed, wing uniformly narrowed from about apical third to base; mouth-edge projecting slightly beyond level of profrons.—European Russia , Siberia; Holarctic.................................... pedella (Fallén) - Axillary lobe of wing developed normally; mouth-edge not projecting beyond level of profrons.—Only Siberia, Russian Far East and China ; Asian................................................................. griseiventris Ringdahl 73. Hind tibia with submedian posterodorsal seta.............................................................. 74 - Hind tibia without submedian posterodorsal seta............................................................ 77 74. Sternite 5 with long and dense hairs; fore tibia with anterodorsal seta in distal half; mid femur in distal third without long posteroventral setae.—European Russia , Siberia; Eurasian................................... trilineella (Zetterstedt) - Sternite 5 with conspicuously shorter hairs; fore tibia without anterodorsal seta in distal half or with a very weak one on fore tibia; mid femur in distal third with strong posteroventrals.................................................... 75 75. Postpedicel long, about 5.0 times as long as wide, reaching mouth-edge; parafacial very narrow, much narrower than width of postpedicel; hind femur with brown dorsal streak; abdomen without distinct spots; arista short pubescent; sternite 5 narrower. Terminalia in Figs 13 A, B .—Caucasus; European............................................ emiliae Lukasheva - With a different combination of characters................................................................ 76 76. Posterodorsal seta on hind tibia as long and strong as anterodorsal seta; arista long-haired, the longest individual hairs as long or a little more than half width of postpedicel; abdomen without spots ( Fig. 12A ).—Siberia, Far East; Asian ( China , Japan )........................................................................................ lacustris Schnabl - Posterodorsal seta on hind tibia half as long as anterodorsal seta or even shorter and hair-like; arista short-haired, the longest individual hairs shorter than half width of postpedicel; abdomen often with dark spots ( Fig. 12C ).—Only Siberia, Far East and China ; Asian........................................................................... luteipes Ringdahl 77. Basal half of hind femur with dense hairs on ventral surface.................................................. 78 - Hind femur without dense hairs on ventral surface.......................................................... 79 78. Fore femur with dark dorsal spot; mid and hind femora with long hairs on anterior-posterior surface; mid femur without preapical anterior seta; sternite 5 with long hairs; abdomen oval and plump.—European Russia ; European................................................................................................. dubiosa Hennig (part) - Fore femur completely yellow; mid femur only with 3 strong posteroventral setae on basal half, with preapical anterior seta; hind femur with short hairs on basal half of ventral surface; sternite 5 without long hairs; abdomen not plump, elongated.— European Russia , Siberia; Eurasian......................................................... perpusilla Meigen 79. All tarsi reddish-yellow, at most middle tarsomeres of fore legs slightly darkened; scutum and scutellum brown dusted; abdomen with more or less distinct paired spots and brown dust between them dorsally; mid femur with a strong anterior and a posterior preapical seta, without posterovental setae at apex, with 1 or 2 weak setae in basal third; mid tibia with antero- and posterodorsal setae the same length and at the same level. Small size: 1.9–2.3 mm .—European Russia , Far East ( Magadan ); Eurasian................................................................................ paludis Tiensuu - With a different combination of characters................................................................ 80 80. Mid femur with a preapical anterior seta; all apical tarsomeres yellowish........................................ 81 - Mid femur without a preapical anterior seta; all apical tarsomeres black......................................... 82 81. Sternite 5 as long as length of sternites 4 + 3, with numerous long and fine hairs ( Fig. 12G ); abdomen without distinct spots; mid femur with 3-4 strong and long anteroventral setae which are longer than diameter of femur; hind femur with long anterovenrals and posteroventrals.—Only Russian Far East................................................. pilipyga Ringdahl - Sternite 5 shorter, as long as sternite 4, without numerous long and fine hairs, only with individual long setae ( Fig. 12H ); abdomen with distinct brown spots; mid femur with 1 or 2 anteroventral setae in basal third; hind femur with short anterovenrals and posteroventrals.—European Russia , Siberia, Far East; Holarctic................................. pumila (Fallén) 82. Mid tibia with anterodorsal seta strong and twice as long as posterodorsal seta.................................... 83 - Mid tibia without anterodorsal seta, but if this seta present it is weak and much shorter than a posterodorsal, the longest anterodorsal and posterodorsal setae equal in length......................................................... 84 83. Pulvilli and claws long; face moderately concave in profile, not long and narrowing below; fore femur often dorsally or even predominantly brown; mid femur with 3–4 long posteroventral setae in basal half. Larger species (4.5–6.0 mm).—European Russia , Siberia; Holarctic..................................................... campestris (Robineau-Desvoidy) - Pulvilli and claws very small; fore femur completely yellow; face not concave in profile, rather long and narrowing below; mid femur without anteroventrals and with 1 long posteroventral seta in basal half. Smaller species (3.0– 3.3 mm ) ( Fig. 10G ). Terminalia in Figs 3 H, I .—Siberia, Far East; Holarctic.................................... conflicta Huckett (part) 84. Mid femur with a row of posteroventral setae, apical seta present; scutum with brown dust between dorsocentral setae, reaching the apex of scutellum. Smaller size: 2.8–3.0 mm............................................................ 85 - Mid femur without posteroventral setae in apical third; scutum and abdomen with different combination of characters. Bigger size: 3.0–5.0 mm..................................................................................... 86 85. Legs yellow; abdomen with brown rectangular marks which merge and seem to form two stripes (Figures in Sorokina 2022 ).— Only North of Russian Far East.......................................................... koni Sorokina (part) - Fore femur with dark dorsal streak, mid and hind femora dark in apical half or third; abdomen with indistinct marks in middle of tergites (Figures in Sorokina 2022 ).—Only North of Russian Far East.......................... gorodkovi Sorokina 86. Postpedicel long, 3.5–4 times as long as wide; scutum without distinct longitudinal stripes; frontal triangle reaching the anterior margin of frons; palpus brownish-yellow on basal half, darker at apex.—Far East ( Kamchatka ); Oriental and Asian.......................................................................................... mandschurica Hennig - Postpedicel not more than 2.5 times as long as wide; scutum with or without distinct longitudinal stripes; frontal triangle reaching or not reaching the anterior margin of frons; palpus brown to black..................................... 87 87. Hind femur with 2–3 strong long anteroventral setae in distal half only and with row of short setae in proximal part; sternite 5 with a wide semicircular median notch, inner edge with numerous short hair-like setae, caudal margin with short setae, pointed ( Fig. 12E ).—Only Siberia and Russian Far East............................................ sibirica Hennig (part) - Hind femur with 3–4 strong long anteroventral setae in basal, middle and apical parts; sternite 5 with a large rectangular median notch, inner edge without numerous short hair-like setae, caudal margin with long setae which are at least as long as sternite 5, rounded ( Fig. 12F )................................................................................... 88 88. Scutum with 3 distinct brown longitudinal stripes. Terminalia in Fig. 13 C, D .—Only Russian Far East and China ; Asian........................................................................................... striolata Hennig - Scutum without distinct longitudinal stripes, or with 2 indistinct stripes......................................... 89 89. Frontal triangle very narrow and whitish grey distally, reaching anterior margin of frons; cercal plate very slender, at least 6.0 times longer than broad. Terminalia in Fig. 13 E, F .—European Russia , West Siberia; Palaearctic........ albicornis Meigen - Frontal triangle gradually narrowed distally, not reaching anterior margin of frons; cercal plate broad, about 3.0 times longer than broad. Terminalia in Fig. 13 G, H .—European Russia , Siberia, Far East; Eurasian............ lineatipes (Zetterstedt)