Unraveling the mysteries of Goeldia Keyserling, 1891: revision, description of seven new species and first record from USA (Araneae: Titanoecidae) Author Almeida-Silva, Lina M. Laboratório de Genômica Evolutiva, Universidade Estadual de Campinas, Instituto de Biologia, Departamento de Genética, Evolução, Microbiologia e Imunologia, Cidade Universitária “ Zeferino Vaz ” Rua Monteiro Lobato, 255, 13083 - 970 - Campinas, SP, Brazil. Author Brescovit, Antonio D. Laboratório de Genômica Evolutiva, Universidade Estadual de Campinas, Instituto de Biologia, Departamento de Genética, Evolução, Microbiologia e Imunologia, Cidade Universitária “ Zeferino Vaz ” Rua Monteiro Lobato, 255, 13083 - 970 - Campinas, SP, Brazil. & Laboratório de Coleções Zoológicas-LCZ, Instituto Butantan, Avenida Vital Brazil, 1500, 05503 - 900, São Paulo, SP, Brazil & Laboratório de Genômica Evolutiva, Universidade Estadual de Campinas, Instituto de Biologia, Departamento de Genética, Evolução, Microbiologia e Imunologia, Cidade Universitária “ Zeferino Vaz ” Rua Monteiro Lobato, 255, 13083 - 970 - Campinas, SP, Brazil. text Zootaxa 2024 2024-03-21 5428 2 151 193 http://dx.doi.org/10.11646/zootaxa.5428.2.1 journal article 10.11646/zootaxa.5428.2.1 1175-5326 10847364 E438D8DA-B142-4A0C-BA25-F6B6EC45C197 Goeldia utcuyacu sp. nov. ( Figs 21 ; 22 ; 25 ) Type material . Holotype ♂ from Utcuyacu ( 11°11’11.9”S 75°27’51.4”W ), Junin , Peru , elev. 1600–2200 m , III.1984 , F. Woytkowski coll. ( AMNH ); paratype female with same data as holotype ( AMNH ). Additional Material Examined. PERU . Cuzco . Ocongate ( 13°37’45”S 71°23’10”W ), Hacienda Capana , elev. 3670 m , 2♀ , 6–11.IV.1947 , J. C. Pallister coll. ( AMNH ) ; Urubamba , Machu-Picchu , ( 13°09’47”S 72°32’44”W ), elev. 2600–2800 m , 1♀ , 1–5.VIII.1964 , B. Malkin coll. ( AMNH ) . Junin : San Ramon ( 11°07’29”S 75°21’25”W ), Estância Naranjal , 2♀ , 20– 7.VII.1965 ( AMNH ) ; Huacapistana ( 11°13’59”S 75°28’59”W ), elev. 1800 m , 1♂ , 2♀ 1juv. , 27–30.VII.1965 ( AMNH ) all collected by P. & B. Wygodzinsky ; San José de Utcuyacu ( 11°11’10”S 75°27’47”W ), 3♀ , 1♂ , 4 juv. , III.1948 , F. Woytkowski coll. ( AMNH ) ; Colonia Perene , Rio Perene , 18 mi. NE La Merced , 2♂ 2♀ , 3.I.1955 , E. Schlinger & E. Ross coll. ( CAS 9021317 , SEM, male palp and female leg IV) . Etymology . The specific epithet is a noun in apposition from the type locality, San José de Utcuyacu, Peru . Diagnosis . Goeldia utcuyacu sp. nov. differs from other species of the genus by the thick patellar apophysis with sinuous tip ( Figs 21B–D ; 22E ); and by the tegular process subrectangular and with enlarged base in the male palp ( Figs 21A ; 22A ). Females can be recognized by the epigynum with median field wide, with posterior edge almost straight ( Fig. 21E ), and the vertical spermathecae slender and longer than copulatory ducts ( Fig. 21F ). FIGURE 21. A–F. Goeldia utcuyacu sp. nov. A–D. Male left palp; E–F Female epigynum: A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view. Description . Male ( Holotype , AMNH). Carapace brown with cephalic area darker; femur I brown; other leg segments yellowish-brown; chelicerae dark brown, labium and endites brown. Total length 4.8. Carapace: length 2.4; width 1.8; height 1.34. Clypeus: height 0.16. Opisthosoma: length 2.43; width 1.4. Eye diameters and interdistances: AME 0.06; ALE 0.11; PME 0.10; PLE 0.12; AME–AME 0.11; AME–ALE 0.11; ALE–PLE 0.5; PME–PLE 0.20; PME–PME 0.13; AME–PME 0.12. Chelicerae paturon: length 1.25. Leg measurements: I: femur 2.2/ patella 1.3/ tibia 2.3/ metatarsus 2.2/ tarsus 1.0/total 9.0; II: 2.0/ 0.8/ 1.9/ 1.85/ 0.95/ 7.5; III: 1.85/ 0.7/ 1.5/ 1.6/ 0.7/ 6.35; IV: 2.35/ 0.7/ 2.2/ 2.0/ 0.85/ 8.1. Leg formula: 1423. Leg spination: femur I–II p1ap; tibia II v2-0-2 or v0-0-2, p1ap, r1ap; III v2sa or v1-0-2, r 0-0-1, p0-1-1; IV: v1-0-2 or v0-1-2, p1ap, r1ap; metatarsus I v1p-2- 1m , p1ap, r1ap; II v2-1r-2(1p-1r)-2(1p-1r)-2ap, p0-2-2, r2ap. Patellar apophysis dorso-posteriorly projected ( Figs 21B–D , 22E ). MLT base enlarged and apex partially sclerotized ( Figs 21C , 22D–E ). Cymbium with macrosetae ( Figs 21B , 22D ). Tegular process with uniform width and base membranous ( Figs 21A ; 22A ). Pars pendula thickening far from embolus tip ( Figs 21D ; 22A ). FIGURE 22. A–E. Goeldia utcuyacu sp. nov. , from Junin, Peru, male genitalia. A. Left palp, bulb, apical view; B. Ditto, retrolateral view; C. Left tibia, ventro-prolateral view; D. Left tibia and cymbium, dorsum apical view; E. Left tibia, dorsum prolateral view. Female (AMNH). Coloration: carapace, chelicerae, and femora dark brown; other leg segments, labium and endites caramel brown. Total length: 6.8. Carapace: length 2.56; width 1.92, height 1.28. Clypeus: height 0.21. Opisthosoma: length 4.4, width 3.2. Eye diameters and interdistances: AME 0.08; ALE 0.13; PME 0.12; PLE 0.13; AME–AME 0.08; AME–ALE 0.13;ALE–PLE 0.04; PME–PLE 0.21; PME–PME 0.14; AME–PME 0.09. Chelicerae paturon: length 1.29. Leg measurements: I: femur 1.85/ patella 0.85/ tibia 1.8/ metatarsus 1.55/ tarsus 0.8/ total 6.85; II: 1.8/ 0.9/ 1.5/ 1.4/ 0.7/ 6.3; III: 1.75/ 0.75/ 1.25/ 1.2/ 0.55/ 5.5; IV: 1.8/ 0.8/ 1.65/ 1.55/ 0.55/ 6.35. Leg formula 1423. Leg spination: femur I–II p1ap; tibia III p1ap; metatarsus I–II v2-2- 1m , p 1ap, r1ap; III v1r-1p-2ap, p1-0-1ap, r0-0-1p; IV v1p-1p- 2m , p1ap, r1ap. Median field of epigynum wide but not closing copulatory openings ( Fig. 21E ). Copulatory duct wall visible by transparency on the median field and partially visible at the copulatory opening area ( Fig. 21E ). Copulatory duct base enlarged ( Fig. 21E–F ). Vertical spermathecae straight ( Fig. 21F ). Horizontal spermathecae straight shorter than vertical spermathecae ( Fig. 21F ). Fertilization ducts enlarged and separated by their length ( Fig. 21F ) Variation . Length ( 2 ♂ ): total 4.20–4.80; carapace 2.00–2.40; femur I 2.10–2.20; ( 10 ♀ ): total 5.20–7.10; carapace 2.14–2.60; femur I 1.78–1.88. Distribution . Peru ( Fig. 25 ).