Unraveling the mysteries of Goeldia Keyserling, 1891: revision, description of seven new species and first record from USA (Araneae: Titanoecidae)
Author
Almeida-Silva, Lina M.
Laboratório de Genômica Evolutiva, Universidade Estadual de Campinas, Instituto de Biologia, Departamento de Genética, Evolução, Microbiologia e Imunologia, Cidade Universitária “ Zeferino Vaz ” Rua Monteiro Lobato, 255, 13083 - 970 - Campinas, SP, Brazil.
Author
Brescovit, Antonio D.
Laboratório de Genômica Evolutiva, Universidade Estadual de Campinas, Instituto de Biologia, Departamento de Genética, Evolução, Microbiologia e Imunologia, Cidade Universitária “ Zeferino Vaz ” Rua Monteiro Lobato, 255, 13083 - 970 - Campinas, SP, Brazil. & Laboratório de Coleções Zoológicas-LCZ, Instituto Butantan, Avenida Vital Brazil, 1500, 05503 - 900, São Paulo, SP, Brazil & Laboratório de Genômica Evolutiva, Universidade Estadual de Campinas, Instituto de Biologia, Departamento de Genética, Evolução, Microbiologia e Imunologia, Cidade Universitária “ Zeferino Vaz ” Rua Monteiro Lobato, 255, 13083 - 970 - Campinas, SP, Brazil.
text
Zootaxa
2024
2024-03-21
5428
2
151
193
http://dx.doi.org/10.11646/zootaxa.5428.2.1
journal article
10.11646/zootaxa.5428.2.1
1175-5326
10847364
E438D8DA-B142-4A0C-BA25-F6B6EC45C197
Goeldia luteipes
(
Keyserling, 1891
)
(
Figs 14
;
27
)
Titanoeca luteipes
Keyserling, 1891: 156
, pl. 4, fig. 108 (
holotype
male from
Rio Grande
(
32°2’ 6”S
52°5’56”W
),
Rio Grande do Sul
,
Brazil
, deposited at NHM, examined).
Calleva paupercula
Simon, 1892: 239
(
holotype
juv
♀
. from Cacheuta (
33°06’S
69°07’W
),
Mendoza
,
Argentina
, deposited at MNHN AR807, examined);
Lehtinen 1967
(erroneously placed as syn. of
G. obscura
).
Syn. nov.
Auximus luteipes
:
Petrunkevitch 1911: 106
.
Amaurobius luteipes
:
Mello-Leitão 1919: 467
, 475;
Bonnet 1955: 290
.
Amaurobius cothurnatus
Mello-Leitão, 1943: 102
, fig. 2 (
holotype
male from Córdoba (
31°25’S
64°11’W
),
Argentina
, deposited at MLP 15620, examined by photo);
Lehtinen 1967
(Syn.).
Amaurobius thoracicus
Mello-Leitão, 1945: 232
(
holotype
juv. from
Argentina
(
37°11’0”S
,
67°22’0”W
), deposited at MLP 16345, examined).
Marusik
et al
. 2020: 54
(sp. inquirendae).
Syn. nov.
Goeldia luteipes
:
Lehtinen 1967: 236
;
Leech 1972: 103
;
World Spider Catalog 2023
.
Additional Material Examined
.
VENEZUELA
.
Bolivar
:
Gran Sabana
,
Parupá
(
5°40’59”N
61°34’59”W
),elev.
1500 m
,
2♂
27.VI–10.VII.1987
,
S. J. Peck
coll.(
AMNH
)
.
BRAZIL
.
Amapá
:
Macapá
(
0°02’18.84”N
51°03’59.10”W
),
1♂
,
2–28.I.2004
,
R. A. Da Silva
coll. (
MCTP 0106580
)
.
Paraíba
:
João Pessoa
,
Campus
UFPB (
7°8’22”S
34°50’43”W
),
1♂
, 2005–2006,
P. L.F. Duarte
coll. (
IBSP 62904
)
.
Distrito Federal
:
Brasília
(
15°47’38”S
47°52’58”W
),
1♂
,
1.XII.2000
,
A. J. Santos
coll. (
IBSP 37516
)
;
Área da Marinha
(
16°0’55”S
48°0’47”W
),
1♂
,
29–31.X.1999
,
G. G. Montingelli
coll. (
IBSP 26545
)
;
Reserva Ecológica do Instituto Brasileiro de Geografia
e
Estatística
(
15°56’41”S
47°53’07”W
),
3♂
,
III–IV.2000
,
Briani
coll. (
IBSP 28384
)
.
Minas Gerais
: Belo Horizonte,
Campus Universidade Federal
de
Minas Gerais
(
19°52’19”S
43°57’58”W
),
1♂
, 3.II.02,
E.S.S. Álvares
coll. (
IBSP 73185
)
;
Estação Ecológica da Universidade Federal
de
Minas Gerais
(
19°52’S
43°58’W
),
3♂
,
27.III.2000
,
14♂
,
VII.1999
–
II.2001
,
E.S.S Álvares
&
E.O. Machado
coll. (
IBSP 27234 ‒27235
,
32290
)
;
Jardim Zoológico
(
44°00’32”W
19°51’41”S
),
1♂
,
13.X.2007
,
A.J. Santos
coll. (
UFMG 1024
)
;
Ouro Preto
,
Parque Estadual do Itacolomi
(
20°25’45.9”S
43°28’28.7”W
),
1♂
,
3♀
,
2–4.XI.2007
, A.J.
Santos
et al.
coll. (
UFMG 1025
)
;
Santana do Riacho
,
Parque Nacional Serra do Cipó
,
Portaria Palácio
(
19°15’S
43°31’W
),
1♂
,
9–11.XII.2001
,
E.S.S. Álvares
&
E.O. Machado
coll. (
UFMG 1411
)
;
Santana do Riacho
,
Parque Nacional da Serra do Cipó
(
19°22’1”S
43°32’17”W
),
5♂
,
3♀
, 2003,
Equipe Biota
coll. (
IBSP 91607
; 91609; 91611‒91612)
;
Uberlândia
,
Reserva Ecológica do Panga
(
19°11’10”S
48°23’30”W
),
3♂
,
IV.2005
–
II.2006
,
M.F. Mineo
coll. (
IBSP 97338
,
97340
)
;
Carrancas
(
21°29’16”S
44°38’34”W
), elev.
1200 m
,
1♀
,
X.2008
,
R.A. Zampaulo
coll. (
IBSP 124140
)
.
Espírito Santo
:
Vila Velha
, APA de
Jacarenema
(
20°24’31”S
40°19’35”W
),
1♂
,
12.X.2005
,
T. Sousa
coll. (
IBSP 133270
)
.
Rio de Janeiro
:
Macaé
,
Restinga de Macaé
(
22°22’15”S
41°47’13”W
),
23.VII.1975
,
L. Moojer
coll. (
MNRJ 3216
)
;
Pinheiral
(
Currently Pinheiro
) (
22°30’46”S
44°00’03”W
),
1♂
, (
MNRJ 926
)
;
Nova Iguaçu
(
22°45’37.16”S
43°26’51.82”W
),
1♀
,
1 juv.
,
Blane
coll. (
MNRJ 929
)
.
São Paulo
:
Itirapina
,
Estação Ecológica de Itirapina
(
22°15’S
47°49’W
),
9♂
,
1 juv.
,
4–9.X.2001
,
C. Bertin
coll. (
IBSP 72122–72124
,
72126–72131
)
;
3♂
, 2001–2002,
Equipe Biota
coll. (
IBSP 97315–97317
)
;
10♂
,
24.III– 06.XII.2000
,
G. Machado
coll. (
IBSP 91614–91611
,
91622
)
;
Campinas
(
22°54’21”S
47°3’39”W
),
1♂
,
23.X.1993
(
IBSP 26641
)
;
1♂
,
16.VIII.2006
(
IBSP 79308
)
;
1♂
,
12.I.1997
(
IBSP 26791
)
;
1♂
,
15.I.2002
(
IBSP 35150
)
, all collected by A.J. Santos;
1♂
,
X.1998
,
A.D. Brescovit
coll. (
IBSP 20129
)
;
Campus da Universidade Estadual de Campinas
(
22°49’3”S
47°4’11”W
),
1♂
,
1.II.2005
,
A.J. Santos
coll. (
IBSP 56973
)
;
1♂
,
18.X.2000
,
A.J. Santos
coll. (
IBSP 35131
)
;
São Carlos
,
Campus da Universidade Federal de São Carlos
(
21°58’S
47°51’W
),
2♀
,
1♂
,
VIII.2006
,
M. Rezende
coll. (
IBSP 88008‒88009
)
;
São Paulo
,
Campus da Universidade
de
São Paulo
(
23°33’44”S
46°43’39”W
),
1♂
,
VIII.1972
,
Students
ICB-USP coll. (
IBSP 4103
)
;
1♂
,
VIII.2005
,
G.R.S. Ruiz
coll. (
IBSP 56129
)
;
1♂
,
G.R.S. Ruiz
coll. (
IBSP 59539
)
;
1♂
,
22.IX.2006
,
G.R.S. Ruiz
coll. (
IBSP 72063
)
;
1♂
,
25.IV.2000
,
A.D Brescovit
coll. (
IBSP 27291
)
;
Osasco
(
23°31’58”S
46°47’31”W
),
1♂
,
1♀
,
III.1991
,
J. Bedana
coll. (
IBSP 5049
)
;
Jaboticabal
(
21°15’18”S
48°19’19”W
),
7♂
, III–
IV/2004
,
F.J. Cividanes
coll. (
IBSP 55860
,
55899
,
55887
)
;
Jundiaí
,
Reserva Natural Municipal Serra do Japi
(
23°14’0”S
46°58’0”W
),
1♂
,
III.2007
A.D. Brescovit
coll. (
IBSP 78555
)
.
Paraná
:
Ponta Grossa
(
25°05’42”S
50°09’43”W
),
1♂
, 2008,
I.M Bortolin
coll (
IBSP 145404
)
.
Santa Catarina
:
Rio Negrinho
(
26°15’14”S
49°31’04”W
),
5♂
,
2 juv.
,
M.A. Nieckel
coll. (
IBSP 68721
, I76007)
.
Rio Grande do Sul
:
Porto Alegre
(
30°01’58”S
51°13’48”W
),
1♂
,
3.IX.1983
,
A.D. Brescovit
coll. (
MCN 12664
)
;
Guaíba
(
30°06’50”S
51°19’30”W
),
1♂
,
4.III.1976
,
H. Gastal
&
M.H. Galileo
coll. (
MCN 27230
)
;
Bom Jesus
(
28°40’04”S
50°25’01”W
),
Fazenda Santa Cruz
,
1♀
,
28–30.III.1998
,
A.B. Bonaldo
coll. (
MCN 29267
)
,
Torres
(
29°20’06”S
49°43’37”W
),
1♂
,
29.XII.1980
–
2.I.1981
,
P. Goloboff
coll. (
MACN 16121
)
;
Eldorado do Sul
(
29°59’49.7”S
51°18’28.3”W
),
2♂
,
25–30.XI.1995
,
G. Carvalho
&
R. Silva
coll (
MCTP 16528
,
12560
)
.
BOLIVIA
:
Cochabamba
:
Cochabamba
(
17°23’37”S
66°9’25”W
),
1♀
, 1940,
J. Vellard
coll. (
MACN 16090
)
;
Monte Puncu
(
17°31’59”S
65°13’59”W
), elev.
2700 m
,
2♀
,
22.X.1983
,
A. Roig.
coll. (
MACN 16024
)
.
ARGENTINA
.
Tucumán
:
Tucumán
(
27°00’S
65°30’W
), 6.25
Km
N of Trancas
,
1♀
,
13.II.1951
,
Ross
&
Michaelbacher
coll. (
CAS 9025978
)
.
Santiago del Estero
:
Moreno
,
Weisburd
(
27°18’S
62°36’W
),
1♀
,
26.IX.1947
(
MACN 16110
)
.
Misiones
:
Misiones
,
Santa Maria
(
27°54’00”S
55°23’00”W
),
1♀
,
XI–XII.1952
,
J. Viana
coll (
MACN 16099
)
;
Chaco
:
Chaco
(
27°27’05”S
58°59’12”W
)
1♀
, 1946,
G. Pinedo
coll. (
MACN 1735
)
.
Córdoba
:
Córdoba
(
31°25’00”S
64°11’00”W
),
2♀
, (
MNRJ 686
)
;
1♀
,
J. Viana
coll. (
MACN 16103
)
;
12♀
,
II.1952
,
J. Viana
coll. (
MACN 16033
)
;
Copina
(
31°33’S
64°42’W
),
2♀
,
2♂
,
XII.1983
,
B. R. de Fernandes
coll. (
MACN 16047
)
;
Cerro Colorado
(
30°05’49”S
63°55’52”W
),
14.X.1961
,
A. de Ferraris
coll. (
AMNH
)
;
Ambúl
(
31°29’20”S
65°03’18”W
),
2♀
,
5.II.1981
.
A. Roig
coll. (
MACN 16128
)
;
Calamuchita
(
31°59’00”S
64°22’00”W
),
El Souce
,
1♀
,
XII.1941
,
J. Viana
coll. (
MACN 2011
)
;
1♀
,
XII.1941
,
M. J. Viana
coll. (
MACN 161183
)
;
Calamuchita
(
31°59’00”S
64°22’00”W
),
4♀
,
1♂
,
I.1955
, (
MACN 16031
)
;
1♀
,
XII.1941
(
MACN 16106
)
;
2♀
,
XII.1941
(
MACN 16104
)
;
2♀
,
III.1957
, (
MACN 16081
)
, all collected by J. M Viana;
Cosquín
(
31°14’00”S
64°27’00”W
),
Camiño
a
la Mandinga
,
1♂
,
10.IX.1990
,
A. Peretti
coll. (
MACN 16049
)
;
Alta Gracia
(
31°40’00”S
64°26’00”W
), 3er paredón,
1♂
,
XI.1985
,
M. E. Galiano
&
F. Miranda
coll. (
MACN 16108
)
;
La Cumbre
(
31°06’00”S
64°28’59”W
),
1♂
,
8.XI.1991
,
M. Ramírez
coll. (
MACN 16043
)
;
Mendoza
:
Mendoza
,
Cacheuta
(
32°53’00”S
68°50’00”W
),
2♀
,
4 juv.
(
MNHN 290
)
.
San Luis
:
Villa Elena
,
Cortaderas
(
32°30’S
65°00’W
),
1♀
,
10.XI.1982
,
E. Maury
coll. (
MACN 16057
)
;
Villa Elena
,
1♂
,
X.1971
,
J. M. Viana
coll. (
MACN 16093
)
.
Buenos Aires
.
Tornquist
,
Sierra de la Ventana
,
Abra de la Ventana
(
38°04’00”S
61°58’59”W
),
3♂
,
2♀
,
19.I.1970
,
M.E. Galiano
coll. (
MACN 16208
)
;
Sierra de la Ventana
(
38°09’S
61°48’W
),
Cerro Negro
,
1♀
,
12.IV.1974
,
C. Cesari
coll. (
MACN 16135
)
;
Tigre
(
34°25’00”S
58°35’00”W
),
1♀
, 1955,
J.M. Viana
coll. (
MACN 16105
)
.
URUGUAY
.
Canelones
:
Marindia
(
34°46’29”S
55°49’13”W
),
3♂
,
16.I.1997
,
Costa
coll. (
FCEN
)
;
Villa
Argentina
(
34°46’00”S
55°47’00”W
),
1♂
,
8.II.1970
,
R. Capocasale
coll. (
CAS 9025953
)
.
Diagnosis
.
Goeldia luteipes
differs from other species of the genus by the narrow, long, and triangular tegular process (
Fig. 14A
) and by the strongly bent tibia in the male palp (
Fig. 14B–C
), and by the black or dark brown color of the proximal half portion of the femora. Females of
G. luteipes
have a wide epigynum that covers the copulatory opening area ventrally (
Fig. 14E
); and copulatory ducts converging anteriorly (
Fig. 14F
).
Description
. Male (IBSP 26545). Coloration: Carapace orange with cephalic area black; palp black; apex of coxa, trochanter, and proximal half of femora black; other leg segments and distal half of femora pale yellow except in leg IV where apex of patella to tarsus black. Opisthosoma black. Total length: 5.64. Carapace: length 2.6, width 1.84, height 1.18. Opisthosoma: length 3.04; width 1.28. Clypeus: height 0.22. Eye diameters and interdistances: AME 0.08; ALE 0.11; PME 0.95; PLE 0.08; AME–AME 0.13; AME–ALE 0.10; ALE–PLE 0.04; PLE–PME 0.19; PME–PME 0.14; AME–PME 0.11. Chelicerae paturon: length 1.41. Leg measurements: I: femur 2.1/ patella 0.72/ tibia 2.02/ metatarsus 1.94/ tarsus 0.92/ total 7.45; II: 1.68/ 0.725/ 1.64/ 2.14/ 0.93/ 7.11; III: 1.76/ 0.72/ 1.5/ 1.4/ 0.63/ 6.01; IV: 2.26/ 0.8/ 2.22/ 1.84/ 0.78/ 7.9. Leg formula: 4123. Leg spination: femur I p1ap, II p1ap; tibia II v1r-2(1r-1p), p1ap, r1ap; III v1-2 or v2sa; r1ap; p1-1ap; IV v1-1-2 or v0-1p-2, p0-1-1ap, r0-0-1ap; metatarsus I v2-1p-2(1r-1p)-2ap or v2-2(1r-1p)-2(1r-1p)-
1m
, r1ap, p1ap; II v2-2(1p-
1m
)-2(1r-
1m
)-
1m
or v2-2-2-
1m
, r1ap; p1ap; d0-2-1p or d0-1p-1p; III v2-2-2-
1m
; p1ap or p1-0; r1ap; d1-2-1r or d0-2-1r; IV v2-0-2-2-2-
1m
, p1-2-0-1ap or p1r-1p-1r-1p-2, r1ap or r0-1r-1, d2ap; tarsus II v2sa; r0-1; III v0-2-2 or v0-1p-2, p0-1; r0-1; IV v0-2-0, r0-2-2 or r0-0-1-1, p0-1-0. Palp with patellar apophysis short and thick (
Fig. 14B–C
); MLT apex partially sclerotized and well developed (
Fig. 14C
).
Pars pendula
thickening subdistal (
Fig. 14D
). Spermatic duct bent over its on axis (
Fig. 14B
).
Female (IBSP 64628). Coloration: carapace brown; coxa to proximal half of the femur brown; distal half of femur and other leg segments pale brown; cephalic area, endites and chelicerae brown, slightly darker than thoracic area; opisthosoma brown. Total length: 5.25. Carapace: length 2.52, width 1.68, height 1.32. Opisthosoma: length 2.94, width 2.16. Clypeus: height 0.15. Eye diameters and interdistances:AME 0.09; ALE 0.14; PME 0.11; PLE 0.1; AME–AME 0.09; AME–ALE 0.11; ALE–PLE 0.05; PLE–PME 0.19; PME–PME 0.18; AME–PME 0.11. Paturon: 1.2 long. Leg measurements: I: femur 1.9/ patella 0.83/ tibia 1.55/ metatarsus 1.45/ tarsus 0.73/ total 6.43; II 1.76/ 0.84/ 1.47/ 1.22/ 0.78/ 6.07; III 1.48/ 0.81/ 1.0/ 0.91/ 0.63/ 4.83; IV 1.85/ 0.8/ 1.56/ 1.32/ 0.64/ 6.17. Leg formula: 1423. Leg spines: absent. Copulatory ducts close together, far from copulatory openings and converging toward middle (
Fig. 14F
). Extra lobe of spermathecae short (
Fig. 14F
)
FIGURE 14.
A–F.
Goeldia luteipes
(
Keyserling, 1891
)
. A–D: male, left palp; E–F: female epigynum: A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view.
Variation
. Length (
10 ♂
): total: 5.20–6.63; carapace: 2.30–3.00; femur I: 1.70–2.88. (
10 ♀
): total 4.10–5.95; carapace 2.05–2.65; femur I 1.38–1.84.
Distribution
.
Venezuela
,
Brazil
,
Bolivia
,
Argentina
, and
Uruguay
(
Fig. 27
).