Unraveling the mysteries of Goeldia Keyserling, 1891: revision, description of seven new species and first record from USA (Araneae: Titanoecidae) Author Almeida-Silva, Lina M. Laboratório de Genômica Evolutiva, Universidade Estadual de Campinas, Instituto de Biologia, Departamento de Genética, Evolução, Microbiologia e Imunologia, Cidade Universitária “ Zeferino Vaz ” Rua Monteiro Lobato, 255, 13083 - 970 - Campinas, SP, Brazil. Author Brescovit, Antonio D. Laboratório de Genômica Evolutiva, Universidade Estadual de Campinas, Instituto de Biologia, Departamento de Genética, Evolução, Microbiologia e Imunologia, Cidade Universitária “ Zeferino Vaz ” Rua Monteiro Lobato, 255, 13083 - 970 - Campinas, SP, Brazil. & Laboratório de Coleções Zoológicas-LCZ, Instituto Butantan, Avenida Vital Brazil, 1500, 05503 - 900, São Paulo, SP, Brazil & Laboratório de Genômica Evolutiva, Universidade Estadual de Campinas, Instituto de Biologia, Departamento de Genética, Evolução, Microbiologia e Imunologia, Cidade Universitária “ Zeferino Vaz ” Rua Monteiro Lobato, 255, 13083 - 970 - Campinas, SP, Brazil. text Zootaxa 2024 2024-03-21 5428 2 151 193 http://dx.doi.org/10.11646/zootaxa.5428.2.1 journal article 10.11646/zootaxa.5428.2.1 1175-5326 10847364 E438D8DA-B142-4A0C-BA25-F6B6EC45C197 Goeldia gauderio sp. nov. ( Figs 11 ; 27 ) Type material. Holotype ♂ from Porto Alegre ( 30°01’58”S 51°13’48”W ), Rio Grande do Sul , Brazil , 11.VII.1963 , C. Hartlieb & Moraes coll., deposited at MCN 1413 ; Paratypes : 1♀ , Porto Alegre ( 30°01’58”S 51°13’48”W ), Rio Grande do Sul , Brazil , 31.VII.1963 , C. Hartlieb & Moraes coll. ( MCN 1002 ) ; 1♀ , 2♂ , 4 juv. from San Luis ( 33°18’S 66°20’W ), 200 Km W of Santa Rosa , San Luis , Argentina , XI.1982 , A. Roig coll. ( MACN 16211 ; IBSP 165960 ) ; 1♀ from Papagayos ( 32°46’1.2”S 65°22’58.8”W ), San Luis , Argentina , 9.IX.1981 , E. Maury coll. ( IBSP 165961 ) . Additional Material Examined. None. Etymology . The specific name is a noun in apposition derived from a regional slang used in the State of Rio Grande do Sul , south of Brazil , generally referring to nomads or vagabond, countryside dwellers. Diagnosis . Males of Goeldia gauderio resemble those of G. luteipes in the dark coloration of the cephalic area and femora. They differ by the position of the spermatic duct median fold at 12 o’clock ( Fig. 11B ), by the thinner patellar apophysis, pointing posteriorly ( Fig. 11C ) and by the short and rectangular tegular process ( Fig. 11A ) in G. gauderio ’s male palp. The female epigynum can be recognized by the fusion of the rim with the copulatory duct wall ( Fig. 11E ). Description. Male ( Holotype ): Coloration: cephalic area and chelicerae reddish brown, thoracic area, endite, labium and sternum brown. Legs with femora completely or almost completely black. Opisthosoma black. Total length 5.15. Carapace: lenght 2.50; width 0.95; height 1.00. Opisthosoma: length 2.6; width 1.7. Clypeus: height 0.15. Eye diameters and interdistances: AME 0.07; ALE 0.11; PME 0.09; PLE 0.09; AME–AME 0.1; AME–ALE 0.11; PME–PME 0.14; PME–PLE 0.23; AME–PME 0.13; ALE–PLE 0.09. Chelicerae paturon: length 1.25. Leg measurements: I: femur 1.88/ patella 0.75/ tibia 1.5/ metatarsus 1.38/ tarsus 0.93/ total 6.43; II: 1.625/0.7/ 1.25/ 1.38/ 0.70/ 5.65; III: 1.45/ 0.7/ 1.25/ 1.15/ 0.68/ 5.23; IV: 1.98/ 1.73/ 0.78/ 1.53/ 0.78/ 6.78. Leg formula 4123. Leg spination femur I–II p1ap; tibia I v1p-2-1r-2, p1-1-1-1, r1ap; II v0-1m-2, r0-1-0-1ap, p0-0-1-1ap; III v1-2; p0-1-0- 1, r1ap; IV v1-0-0-1-2-0, r1ap; metatarsus I v2-1p-1p-1r-1p- 1m , p0-1-0-1ap, r1ap; II v1-2- 1m , r0-1-0-1ap, p1ap; III v2-2- 1m , p1-1-1, r1ap; IV v2-2-2- 1m , p1ap, r1ap. Retrolateral basal side of the tibia and retrolateral apical side of the patella with a knob ( Fig. 11B, D ). Patellar apophysis with a thin depression ( Fig. 11C ). MLT apex partially sclerotized and long ( Fig. 11C ). Tegular process as depicted ( Fig. 11A ). Pars pendula thickening subdistal ( Fig. 11D ). Female ( Paratype , MCN 1002): Coloration: carapace brown with cephalic area and chelicerae slightly darker. Endites, labium and sternum brown. Legs brown with femora slightly darker. Opisthosoma brown. Total length 8.00. Carapace: length 3.28; width 2.9; height 1.6. Opisthosoma 4.08 long; 3.2 wide. Clypeus: height 0.29. Chilum absent. Eye diameters and interdistances: AME 0.1; ALE 0.12; PME 0.1; PLE 0.12; AME–AME 0.13; AME–ALE 0.21; PME–PME 0.22; PME–PLE 0.4; AME–PME 0.17; ALE–PLE 0.15. Chelicerae paturon: length 1.81. Leg measurements: I: femur 2.15/ patella 1.10/ tibia 1.75/ metatarsus 1.69/ tarsus 0.91/ total 7.59; II: 1.6/ 0.97/ 1.47/ 1.56/ 0.81/ 6.41; III: 1.60/ 0.72/ 1.16/ 1.28/ 0.6/ 5.36; IV: 2.25/1.88/1.00/1.56/ 0.68/ 7.37. Leg formula 1423. Leg spination: femur I–II p1ap; tibia III p0-1-1, r1ap; IV v1sap, p 1ap, r1ap; metatarsus I v2-1p-2- 1m , p1ap, r0-1-1; II v2-2- 1m , p1ap, r1ap; v2-1p-2- 1m , p1ap, r1ap, d1p-1p-2ap; III v2-1p-2- 1m , p1ap, r1ap, d1p-1p-2ap; IV v2-1p-1p- 1m , p1ap, r1ap, d2ap; tarsus III v 1m ; IV v2ap. Copulatory duct wall fused to the epigynal rim ( Fig. 11E ). Posterior lateral edge of the epigynum rounded and sinuous ( Fig. 11E ). Copulatory ducts apex dorsally curved ( Fig. 11F ). Extra lobe of the spermathecae L-shaped ( Fig. 11F ). Vertical spermathecae bent horizontally ( Fig. 11F ). Horizontal spermathecae very short and almost round ( Fig. 11F ). Variation . Length ( 3♂ ): total 5.05–5.30; carapace 2.50–2.80; femur I 1.65–1.88. ( 2♀ ): total 5.40–6.60; carapace 2.10–2.70; femur I 1.56–1.82. Distribution . State of Rio Grande do Sul , Brazil and Province of San Luis , Argentina ( Fig. 27 ).