A taxonomical revision of Ilex (Aquifoliaceae) in the Pan-Himalaya and unraveling its distribution patterns
Author
Hong, De-Yuan
text
Phytotaxa
2015
2015-10-08
230
2
151
171
http://dx.doi.org/10.11646/phytotaxa.230.2.3
journal article
10.11646/phytotaxa.230.2.3
1179-3163
13630869
14.
Ilex rotunda
Thunberg (1784: 168)
.
Type
:—
JAPAN
. not designated
Ilex rotunda
was recorded from
Japan
and S
Korea
southwestwards to
Vietnam
, and
Yunnan
,
China
, while
I. excelsa
(Wall.) Voigt (1845: 347)
was reported from N
Pakistan
eastwards to
Yunnan
of
China
via N
India
,
Nepal
, and
Bhutan
. Therefore, both these two species are present in Yunnan.
Hu (1949a)
and
Chen
et al.
(2008)
did not use external morphology to distinguish these two species. Instead, they used only characters of the pyrenes: sulcate vs. esulcate, and endocarp coriaceous vs. woody. The present author examined a large number of specimens of the two species from
Japan
to NW
India
, but also failed in finding any significant difference in external morphology. The result of observations of the pyrenes shows that the pyrenes from
Japan
(two collections),
Hongkong
(one collection), and Guangxi (one collection) are all 3-striate and 2-sulcate, whereas those from Tengchong (SW Yunnan) are esulcate. The specimens from the other parts of Yunnan have a diverse pyrene morphology. The specimens from Pingbian, Yanshan and Xichou in SE Yunnan have pyrenes which are 2-striate and 1-sulcate on the dorsal side, whereas those from Luchuan in C Yunnan and Lincang in SW Yunnan have the pyrenes 2-striate but esulcate on the dorsal side (
Fig. 6
). It is, therefore, reasonable to treat these two groups as two subspecies.
1a. Pyrenes 3-striate and 2-sulcate, rarely 2-striate and 1-sulcate .................................................................
Ilex rotunda
subsp.
rotunda
1b. Pyrenes smooth........................................................................................................................................
Ilex rotunda
subsp.
excelsa