The genus Leucophenga (Diptera, Drosophilidae), part VIII: twenty-one species from the Oriental region, with morphological and molecular evidence Author Huang, Jia Author Wang, Yalian Author O’Grady, Patrick Michael Author Su, Yirui Author Chen, Hongwei hongweic@scau.edu.cn text Zootaxa 2018 2018-10-19 4503 1 1 70 http://dx.doi.org/10.11646/zootaxa.4503.1.1 journal article 10.11646/zootaxa.4503.1.1 1175-5326 5297894 5F058E32-AEF1-4070-8CC8-177B4CE140BA Leucophenga forcipula Huang & Chen , sp. nov. ( Figs. 3E , 7D–F , 10G, 10H , 15C, 15D , 28 ) Diagnosis. This species is most similar to L . hyaloptera sp. nov. in having the pleura with a brown, longitudinal stripe above ( Figs. 7F , 8C ), the wing entirely hyaline ( Figs. 3E , 4A ), the colour patterns of abdominal tergites in females ( Figs. 10H , 11E ), and the shape and pubescence of epandrium, cercus, and surstylus ( Figs. 28A , 32A ), but can be distinguished from the latter in having the aedeagus laterally with a long, distally narrow, apically acute process, which bearing a sensillum at tip ( Fig. 28D ); see below that species. FIGURE 28. Leucophenga forcipula Huang & Chen , sp. nov. , male terminalia in lateral view. A. Epandrium, surstylus, and cercus; B. Hypandrium, paramere, gonopods, aedeagus, and aedeagal apodeme; C. Paramere; D. Aedeagus. Scale bars = 0.1 mm. Description. Ocellar triangle brownish yellow to brown, with ca. 2 setae above ocellar setae. Frons brownish yellow ( Fig. 7D ). Pedicel dark yellow; first flagellomere yellow ( Fig. 7D ). Clypeus entirely brownish yellow. Palpus yellow ( Fig. 7D ). Mesonotum mostly brownish yellow ( Fig. 7E ). Postpronotal lobe yellow ( Fig. 7F ). Acrostichal setulae in ca. 12–14 irregular rows ( Fig. 7E ). Scutellum brownish yellow to brownish, dark brown laterally, yellow at tip ( Fig. 7E ). Katepisternum and mesopleuron mostly yellow ( Fig. 7F ). Wing ( Fig. 3E ): Costal vein between R 2+3 and R 4+5 distally with ca. 4 or 5 peg-like spinules on ventral surface; R 4+5 and M 1 nearly parallel distally; halter mostly yellowish, with a brown to dark brown patch at knob ( Fig. 7E ). Abdominal tergites dark brown to black, 2nd tergite yellow medially and laterally, 3rd tergite mostly yellowish except for lateral region, only setigerous laterally, 5th tergite with yellow patches submedially and sublaterally in males ( Fig. 10G ); 3rd to 5th tergites with yellow patches submedially and sublaterally in females ( Fig. 10H ). Male terminalia: Epandrium with ca. 6 setae near posterior and ventral margins per side ( Fig. 28A ). Hypandrium with 2 sensilla ( Fig. 28B ). Paramere slender, narrow basally and rounded apically, with ca. 6 sensilla distally ( Fig. 28C ). Aedeagus rounded apically ( Fig. 28D ). Measurements and indices. BL = 2.93 mm in holotype (range in 2♀ paratypes : 3.00–3.20), ThL = 1.43 mm (1.47–1.53), WL = 2.60 mm (2.65–2.73), WW = 1.20 mm (1.20–1.23), arb = 7/3 (8/3), avd = 0.81 (0.80–0.83), adf = 2.13 (1.88–2.00), flw = 2.07 (1.63–1.87), FW/HW = 0.34 (0.33–0.36), ch/o = 0.04 (0.05), prorb = 0.076 (0.73– 0.74), rcorb = 0.72 (0.69–0.70), vb = 0.39 (0.38), dcl = 0.38 (damaged and 0.47), presctl = 0.48 (damaged and 0.50), sctl = 1.19 (damaged and 1.35), sterno = 0.75 (damaged and 0.78), orbito = 1.75 (1.82–2.00), dcp = 0.21 (0.21–0.22), sctlp = 0.91 (0.97–1.00), C = 2.35 (2.14–2.19), 4c = 1.21 (1.23–1.24), 4v = 2.12 (1.97–2.06), 5x = 1.50 (1.29–1.31), ac = 2.35 (2.05–2.33), M = 0.73 (0.62–0.63), C3F = 0.73 (0.76–0.77). Type specimens. Holotype ( SCAU , No. 128580), CHINA : Menglun , Mengla , Yunnan , 21°41'N , 101°25'E , alt. 900m , 28.ix.2011 , ex tussock, HW Chen . Paratypes : CHINA : 2♀ ( SCAU , Nos 128581, 82), 780m , 17.iv.2007 , other data same as holotype . Distribution. China ( Yunnan ). Etymology. From the Latin word “ forceps ” (= forcipate), referring to the shape of the process on aedeagus in lateral view.