First record of Mythicomyia Coquillett, 1983 from Colombia with description of a new species of Mythicomyia (Heterhybos) Brèthes, 1919 (Diptera: Mythicomyiidae: Mythicomyiinae) and an updated key to South American species Author Ramos-Pastrana, Yardany Universidad de la Amazonia, Grupo de Investigación en Entomología Universidad de la Amazonia - GIEUA-, Laboratorio de Entomología - LEUA-, Av. 11 5 - 69 Juan XXIII, Florencia, Caquetá, Colombia Author Rojas-Portilla, Edilberto 0000-0002-1314-755X Universidad de la Amazonia, Grupo de Investigación en Entomología Universidad de la Amazonia - GIEUA-, Laboratorio de Entomología - LEUA-, Av. 11 5 - 69 Juan XXIII, Florencia, Caquetá, Colombia neale@bishopmuseum.org text Zootaxa 2024 2024-01-08 5397 4 539 550 http://zoobank.org/a3f91a4f-f99b-4dbc-914a-3542b33fcc03 journal article 284798 10.11646/zootaxa.5397.4.5 359e2512-66e8-4ff5-9dfc-b1d621f6c30e 1175-5326 10469180 A3F91A4F-F99B-4DBC-914A-3542B33FCC03 Mythicomyia Coquillett, 1893 Mythicomyia Coquillett, 1893: 209 (in Empididae ). Williston 1896: 73 (in Empididae ). Melander, 1902: 337 (in Empididae ). Aldrich, 1905: 218 (in Leptidae ). Cresson, 1915: 449 (in Empididae ). Greene, 1924: 60 . Melander, 1961: 161 . Painter & Painter, 1965: 416 . Cole & Schlinger, 1969: 238 . Hull, 1973: 266 . Painter, Painter & Hall, 1978: 11 . Hall & Evenhuis, 1986: 327 . Hall & Evenhuis, 1987: 596 . Evenhuis, 2002a: 36 . Evenhuis, 2022: 1 . Type species: Mythicomyia riley Coquillett (1893) , by monotypy. Diagnosis. [adapted from Hall (1976) ]. Male holoptic; female dichoptic. Body destitute of macrochaetae. Head globular, attached by a distinct cervical sclerite; postcranium not produced; antennae with three segments, scape very short, nearly indiscernible, pedicel as long as wide, two flagellomeres and stylus placed subapically on the second flagellomere; proboscis short or long, almost always projecting beyond oral margin. Thorax humpbacked, higher than long, legs slender, tibiae at most with small apical spurs, pulvilli well developed, distinct; wing with costal vein incomplete, ending just beyond end of vein R 4+5 , cell dm much longer than wide or open apically, well-developed keel-shaped anal lobe, vein separating basal cells not evident. Abdomen round, somewhat pointed with seven visible segments in both sexes. Males with or without secondary sexual character modifications on the legs (middle tibia and basitarsus of male of the nominate subgenus often contorted or, with modified shapes of and notches in mid and/or hind legs and presence of pegs and large setae). Male genitalia with epandrium large, ejaculatory apodeme with apex acute or truncated, lateral arms of parameral sheath thin with apex truncated or acute, gonocoxa simple or with lobe, gonostylus downward-directed, in the opposite direction to the epandrium.