Qorimayus, a new genus of relictual, high-altitude harvestmen from western Argentina (Arachnida, Opiliones, Gonyleptidae) reveals trans-Andean phylogenetic links Author Acosta, Luis E. text Zootaxa 2020 2020-01-13 4722 2 129 156 journal article 24373 10.11646/zootaxa.4722.2.2 c31e76f5-34bc-48e9-91d2-e0f225ebda09 1175-5326 3605827 1206855A-A34A-474A-AA5B-D1656C418703 Metabalta polyhastata ( Hara, 2016 ) comb. nov. Nanophareus polyhastatus Hara, 2016: 117 ; Pérez-Schultheiss et al., 2019: 10 . urn:lsid:zoobank.org:act: 364BA304-56DD- 4C73-9BF3-FA652B038A46 The results of the cladistic analysis ( Fig. 1 ) demonstrated that Nanophareus polyhastatus Hara, 2016 forms a wellsupported clade with species of Metabalta . This relationship is underpinned α-taxonomically too, as evidenced by a close examination of the original description and illustrations of N. polyhastatus : the exomorphological and genital features of this species ( Hara, 2016 ) best match the character states present in Metabalta , as defined in Table 8 . For example, fig. 6A by Hara (2016) clearly displays pedipalp coxae with ‘normal’ appearance, i.e., not elongated as in Nanophareus . In addition, N. polyhastatus has a ‘normal’ development of basichelicerite and pedipap femur, and the retrolateral apical-subapical spines on tibia are placed in a bifid basal socket, not as large and furcate-tipped as in Nanophareus ( Table 8 , chars. b, d, h). Several genital characters support the transfer of N. polyhastatus into Metabalta , like the sigmoid curvature of stylus, its orientation and apical narrowing, and the apical end of VPS ( Table 8 , chars. o, p, r, s); drawings of Hara (2016) are not detailed enough to see if also char. t (subapical spines on VPS shaft) applies. Based on this evidence, I hereby formally propose the new combination Metabalta polyhastata ( Hara, 2016 ) comb. nov . (the original spelling polyhastatus changed to polyhastata to ensure the agreement in grammatical gender with the genus name Metabalta ). Aside, Pérez-Schultheiss et al. (2019) suggested that N. polyhastatus and Metabalta albipes Mello-Leitão, 1931 might prove to be the same. Although the latter species has no original figure available for a comparison, that of Metabalta porteri Mello-Leitão, 1936 (currently under synonymy of M. albipes , after Ringuelet 1959 ) looks identical to M. polyhastata , as described and drawn by Hara (2016) ; key similarities refer to the general apophysis pattern on femur IV, and the shape (and thickness) of apophysis of coxa IV. Until the relevant types are studied I prefer not to formalize this presumed specific synonymy. It should be noted that M. polyhastata was included by Hara (2016) in his cladistic analysis of Nanophareus , but resulted in an internal position in the genus. Additional material examined. Metabalta efformata : Chile : Sierra de Chillán , 2 ♂ syntypes ( SMF RII 952 ) ; Metabalta geniculata : Chile : Valparaíso , 1 ♂ , 1 ♀ syntypes ( SMF RII 951 ) ; Nanophareus palpalis : ‘ Chile’ , 1 ♂ lectotype , 1 ♂ , 2 ♀ paralectotypes (designated by Hara et al. 2012 ; SMF 986 /1); Nanophareus bosqenublado : Chile : Los Molles , elev. 2 m , under succulent rock cover along coast, 9 Jan. 1985 , N.I. Platnick & O.F. Francke ( AMNH ) .