Five new species of lapsiine jumping spiders from Ecuador (Araneae: Salticidae)
Author
Maddison, Wayne P.
text
Zootaxa
2012
3424
51
65
journal article
10.5281/zenodo.208849
f95e4c61-f1a8-4333-a598-b20d703e56d2
1175-5326
208849
Lapsias lorax
,
sp. nov.
(
Figs 2–9
)
Type
material.
Holotype
male in
QCAZ
, temporarily held at the UBC-SEM, with data: "
ECUADOR
: Pichincha:
Bellavista
Cloud Forest Reserve, Discovery Trail.
S 0.01462
W 78.68326
.
2240 m
elev.
9 November 2010
. W Maddison. WPM#10-067", "Male W624 Photo'd
9 Nov
#
ECU
2010-3221", "UBC-SEM AR00194"
Etymology.
Named after Theodor Geisel's fictional character The
Lorax
, a small creature with yellow mustache who advocated for the preservation of trees. This name was suggested by
Tristan
Long to refer to the bars of yellow setae on the chelicerae (appearing as a mustache) and to reflect on the importance of preserving forests such as those where this spider was found.
Diagnosis.
The placement of the embolus, conductor and median apophysis toward the distal tip of the palp is more extreme than in other
Lapsias
. Accordingly, the straight portion of the sperm duct along the prolateral margin of the tegulum stands out as being particularly long, before the duct turns retrolaterally to meet the median apophysis. The yellow bars of setae across the chelicerae appear to be unique among
Lapsias
, although Simon's specimens of the other species are old and there is the possibility that the setae may have been lost (
type
series of
L. estebanensis
,
L. tovarensis
,
L. ciliatus
, and
L. cyrboides
, in MNHN Paris examined).
Notes.
Two possible synapomorphies with the
type
species,
Lapsias estebanensis
, could be proposed as justification for placing this new species in the genus. The first is the very short embolus (shared also with
L. tovarensis
, but also with the genus
Soesiladeepakius
); the second is the migration of the retrolateral tibial apophysis dorsally to approach a second dorsal apophysis (
Figs. 9–10
). In
L. estebanensis
,
L. tovarensis
, and
L. lorax
, a dorsal view of the palp tibia shows a V-shaped cleft framed by two ridges, the lateral representing the retrolateral tibial apophysis, and the medial representing a dorsal apophysis. This cleft is not apparent in
L. ciliatus
or
L. cyrboides
. In body form and markings,
L. lorax
is quite similar to
L. tovarensis
.
Description.
Male
(
holotype
). Carapace length 2.0; abdomen length 2.1. PME small. Chelicerae not particularly enlarged, with 4 promarginal and 2 retromarginal teeth, closely placed (more or less fissident). Palpus (
Figs 7–9
) with short terminal embolus and small, thin median apophysis which is hooked at the tip. Accompanying the median apophysis is a membranous fold that probably is the conductor. Tibial apophysis short, shifted dorsally. Tibia of first leg with 3 pairs of ventral macrosetae; first metatarsus with 4 prolateral and three retrolateral macrosetae. Colour (
Figs 2–6
): honey-coloured (in alcohol, honey-cream) with dark brown markings. Thoracic area dark except two pale side bars and a central pale stripe; ocular quadrangle brown. Clypeus pale, with white setae. Chelicerae black in front, with a distinct oblique yellow band of setae just below the clypeus (
Fig. 2
). Femur of palp mostly pale; patella, tibia and cymbium dark. Femora of legs pale except dark spots dorsally. Patellae and tibiae orange-brown. Metatarsi and tarsi with dark termini, especially on legs 3 and 4. Abdomen medium brown, with chevrons.
Natural history.
The single male was found by shaking lianas climbing within moss covering a tree trunk in cloud forest. Extensive subsequent searching for one day in this microhabitat yielded many
Thrandina bellavista
(
4 adults
and
23 juveniles
), but no more specimens of
Lapsias lorax
. This may indicate the species is uncommon, or that the location where it was found was not its core habitat—it may, for instance, live primarily in the canopy or at higher elevation. When walking, the second pair of legs is waved periodically, as seen in
Thrandina
and many other basal salticids (Maddison, 2006, 2009). A video of the living
holotype
is available at http://www.youtube.com/watch?v=Tgf2JCdr_88.