<document id="E6970D0C60A245C271827503C6430C6D" ID-DOI="10.11646/zootaxa.5326.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="8243811" ID-ZooBank="2EC25BF0-D36F-4029-AD1C-A9B62A668FEE" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1691944079938" checkinUser="plazi" docAuthor="Werner, Maria Johanna, Hausmann, Axel, Kostjuk, Igor, Wanke, Dominic &amp; Rajaei, Hossein" docDate="2023" docId="A420BA7FF059FFE5FF1EFC447F9BFAD6" docLanguage="en" docName="zootaxa.5326.1.1.pdf" docOrigin="Zootaxa 5326 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5326.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Phaselia erika Ebert 1965" docType="treatment" docVersion="5" lastPageNumber="17" masterDocId="5819C207F049FFF5FF89FFA27F76FFE3" masterDocTitle="Integrative taxonomic revision of the genus Phaselia Guenée, [1858] (Geometridae: Ennominae) in the Middle East and Central Asia" masterLastPageNumber="66" masterPageNumber="1" pageNumber="17" updateTime="1699557214275" updateUser="plazi">
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<mods:title id="F16A1E859CAF1C2F1215C289C4405D74">Integrative taxonomic revision of the genus Phaselia Guenée, [1858] (Geometridae: Ennominae) in the Middle East and Central Asia</mods:title>
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<mods:namePart id="203CB21FF6987FEDA3899A4D34BE47A4">Werner, Maria Johanna</mods:namePart>
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<mods:namePart id="5AE70C1B0ABE0E8080854F4F65721F1B">Hausmann, Axel</mods:namePart>
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<taxonomicName id="EB8970EAF059FFE5FF1EFC447EABFBE3" ID-CoL="4FVZ3" ID-ENA="3045205" authority="Ebert, 1965" authorityName="Ebert" authorityYear="1965" box="[151,477,998,1025]" class="Insecta" family="Geometridae" genus="Phaselia" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="erika">
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<emphasis id="1EFDD77BF059FFE5FF1EFC447E33FBE2" bold="true" box="[151,325,998,1025]" italics="true" pageId="16" pageNumber="17">Phaselia erika</emphasis>
<bibRefCitation id="48187698F059FFE5FEC5FC457EABFBE3" author="Ebert, G." box="[332,477,998,1025]" pageId="16" pageNumber="17" pagination="1 - 32" refId="ref30159" refString="Ebert, G. (1965) Afghanische Geometriden (Lep.) I. Stuttgarter Beitrage zur Naturkunde, 142, 1 - 32." type="journal article" year="1965">Ebert, 1965</bibRefCitation>
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<subSubSection id="649358E2F059FFE5FF1EFBA97D14FBC6" box="[151,610,1034,1061]" pageId="16" pageNumber="17" type="description">
<paragraph id="2C360B69F059FFE5FF1EFBA97D14FBC6" blockId="16.[151,610,998,1061]" box="[151,610,1034,1061]" pageId="16" pageNumber="17">
(
<figureCitation id="B4B217ECF059FFE5FF16FBA97E57FBC6" box="[159,289,1035,1061]" captionStart="FIGURES 33–58" captionStartId="30.[151,264,220,244]" captionText="FIGURES 33–58: Wing pattern and coloration of Phaselia species. 33–36: P. erika erika. 33: Holotype, ♁ (Afghanistan, Sarobi, g. prep. G25 G. Ebert); 34–35: Paratype, ♀ (Afghanistan, Sarobi); 36: ♀ (West Pakistan, Salt Range, Chingi, g. prep. 0090/2021 M. Werner); 37–40: P. erika jonubi ssp. nov. 37: Holotype, ♁ (S Iran, Hormozgan prov., Beshgagerd Mts., Davari vil., g. prep. 0094/2021 M. Werner); 38: Paratype, ♀ (Iran, Bandar-Abbas, g. prep. 0016/2021 M. Werner); 39: Paratype, ♀ (Iran, Balutsch- estan, Khasch, 11 km NE Karvandar, g. prep. 0045/2021 M. Werner); 40: Paratype, ♁ (Iran, Djiroft, Anbar Abad, g. prep. 0070/2021 M. Werner); 41–42: P. erika sindhi ssp. nov. 41: Holotype, ♀ (Pakistan, Sindh, Thar, g. prep. 0136/2021 M. Werner); 42: Paratype, ♀ (Pakistan, Sindh, Mirpurkhas, g. prep. 0135/2021 M. Werner); 43–46: P. sihvoneni sp. nov. 43: Holotype, ♀ (India, Ladakh Kharbu, g. prep. 1213/2021 D. Wanke); 44: Paratype, ♁ (N Pakistan, N Sost, g. prep. 0159/2021 M. Werner); 45: Paratype, ♁ (N Pakistan, 2 km N of Karimabad, g. prep. 0153/2021 M. Werner); 46: Paratype, ♁ (Tajikistan, Rushan, Shipad, g. prep. 0130/2021 M. Werner); 47–49: P. kasyi. 47: Holotype, ♁ (Afghanistan, Band-i-Amir, g. prep. 15.911 Mus. Vind.); 48: ♁ (Afghanistan, Band-i-Amir, g. prep. 0061/2021 M. Werner); 49: ♀ (Afghanistan, Khindjan, g. prep. 0042/2021 M. Werner); 50–53: P. deliciosaria. 50: ♁ (Turkey, Belen pass, g. prep. 0027/2021 M. Werner); 51: ♁ (Turkey, Urfa, g. prep. 0155/2021 M. Werner); 52: ♁ (Syria, Akbes [Meydan Ekbaz], g. prep. 0163/2021 M. Werner); 53: ♁ (Israel, Nahal Amud National Reserve, g. prep. 0114/2021 M. Werner); 54–55: P. sp. cf. deliciosaria 54: ♁ (Oman, Al Mughsayl, g. prep. 0150/2021 M. Werner); 55: ♀ (Oman, Al Mughsayl, g. prep. 0149/2021 M. Werner); 56–58: P. joestleinae. 56: Holotype, ♁ (Turkey, Adana, g. prep 5214 ZSM); 57: Paratype, ♀ (Turkey, Adana, g. prep 8560 ZSM); 58: ♀ (Turkey, Döşemealti Çubuk boěazi, g. prep. 0026/2021 M. Werner). a = upside; b = underside." pageId="16" pageNumber="17">Figs 33–42</figureCitation>
, 
<figureCitation id="B4B217ECF059FFE5FEA5FBA97E03FBC6" box="[300,373,1035,1061]" captionStart-0="FIGURES 86–90" captionStart-1="FIGURES 91–97" captionStartId-0="36.[151,264,1908,1932]" captionStartId-1="37.[151,264,1821,1845]" captionTargetBox-0="[174,1414,186,1870]" captionTargetBox-1="[206,1381,187,1793]" captionTargetId-0="figure-26@36.[168,1419,181,1885]" captionTargetId-1="figure-18@37.[201,1386,181,1798]" captionTargetPageId-0="36" captionTargetPageId-1="37" captionText-0="FIGURES 86–90: Variation of the male genitalia of Phaselia erika erika. 86: Holotype, from: Afghanistan, Sarobi (g. prep. 25 G. Ebert); 87: from: N Pakistan, Shandur Pass (g. prep. 0152/2021 M. Werner); 88: from: Afghanistan, Sarobi, g. prep. 0138/2021 M. Werner); 89: from: Afghanistan, Sarobi (g. prep. 0139/2021 M. Werner); 90: from: Pakistan, Chingi (g. prep. 0089/2021 M. Werner). a = genitalia capsule; b = aedeagus. Scalebar = 1 mm." captionText-1="FIGURES 91–97: Variation of the male genitalia of Phaselia erika jonubi ssp. nov. 91: Holotype, from: Iran, Hormozgan, Davari (g. prep. 0094/2021 M. Werner); 92: Paratype, from: Iran, Baloutchistan, Bender Tchahbahar (g. prep. 0044/2021 M. Werner); 93: Paratype, from: Iran, Fars, S Zagros (g. prep 0055/2021 M. Werner); 94: Paratype, from: N-Oman, Musandam, Aqabat Oso (g. prep. 0093/2021 M. Werner); 95: Paratype, from: Iran, Djiroft, Anbar Abad (g. prep. 0070/2021 M. Werner); 96: Paratype, from: Iran, Esfahan, NE Anarak (g. prep. 0050/2021 M. Werner); 97: Paratype, from: S Iran, Bandar-Abbas (g. prep. 0017/2021 M. Werner). a = genitalia capsule; b = aedeagus. Scalebar = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.8243831" figureDoi-1="http://doi.org/10.5281/zenodo.8243835" httpUri-0="https://zenodo.org/record/8243831/files/figure.png" httpUri-1="https://zenodo.org/record/8243835/files/figure.png" pageId="16" pageNumber="17">86–97</figureCitation>
, 
<figureCitation id="B4B217ECF059FFE5FE09FBA87E92FBC6" box="[384,484,1034,1061]" captionStart-0="FIGURES 128–130" captionStart-1="FIGURES 131–135" captionStartId-0="46.[151,264,1943,1967]" captionStartId-1="47.[151,264,1856,1880]" captionTargetBox-0="[161,1427,188,1914]" captionTargetBox-1="[237,1350,182,1827]" captionTargetId-0="figure-26@46.[155,1432,181,1919]" captionTargetId-1="figure-18@47.[232,1355,181,1832]" captionTargetPageId-0="46" captionTargetPageId-1="47" captionText-0="FIGURES 128–130: Female genitalia of Phaselia erika erika. 128: Paratype, from:Afghanistan, Sarobi (g. prep. 24 G. Ebert); 129: from: Afghanistan, Nuristan (g. prep. 1199/2021 D. Wanke); 130: from: Afghanistan, Asmar, Kunartal (g. prep 0063/2021 M. Werner). Scalebar = 1 mm." captionText-1="FIGURES 131–135: Female genitalia of Phaselia erika ssp. 131–133: P. erika jonubi ssp. nov. 131: Paratype, from: Iran, Kerman, Baft, Khabr (g. prep. 0164/2021 M. Werner); 132: Paratype, from: Iran, Baluchestan, Khasch, NE Karvandar (g. prep. 0045/2021 M. Werner); 133: Paratype, from: Iran, Bandar-Abbas (g. prep. 0016/2021 M. Werner); 134–135: P. erika sindhi ssp. nov. 134: Holotype, from: Pakistan, Thar (g. prep. 0136/2021 M. Werner); 135: Paratype, from: Pakistan, Sindh, Mirpurkhas (g. prep. 0135/2021 M. Werner). Scalebar = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.8243855" figureDoi-1="http://doi.org/10.5281/zenodo.8243857" httpUri-0="https://zenodo.org/record/8243855/files/figure.png" httpUri-1="https://zenodo.org/record/8243857/files/figure.png" pageId="16" pageNumber="17">128–135</figureCitation>
, 
<figureCitation id="B4B217ECF059FFE5FE79FBA97D6AFBC6" box="[496,540,1035,1061]" captionStart="FIGURE 144" captionStartId="50.[151,250,884,908]" captionTargetBox="[157,1432,191,859]" captionTargetId="figure-26@50.[151,1435,181,861]" captionTargetPageId="50" captionText="FIGURE 144: Distribution pattern of Phaselia species in Middle East and Central Asia." figureDoi="http://doi.org/10.5281/zenodo.8243865" httpUri="https://zenodo.org/record/8243865/files/figure.png" pageId="16" pageNumber="17">144</figureCitation>
, 
<figureCitation id="B4B217ECF059FFE5FDA1FBA87D20FBC6" box="[552,598,1034,1061]" captionStart="FIGURE 145" captionStartId="51.[151,250,1941,1965]" captionTargetBox="[264,1307,184,1898]" captionTargetId="figure-18@51.[253,1334,179,1903]" captionTargetPageId="51" captionText="FIGURE 145: Maximum likelihood analysis of 70 species of the genus Phaselia, based on COI 5’ sequences (built with MEGA X; Kimura 2-parameter model; bootstrap method, 1000 replications using MEGA X (Kumar et al., 2018))." figureDoi="http://doi.org/10.5281/zenodo.8243869" httpUri="https://zenodo.org/record/8243869/files/figure.png" pageId="16" pageNumber="17">145</figureCitation>
).
</paragraph>
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<subSubSection id="649358E2F059FFE5FF1EFBED7F9BFAD6" pageId="16" pageNumber="17" type="reference_group">
<paragraph id="2C360B69F059FFE5FF1EFBED7EDCFB65" blockId="16.[151,1436,1103,1158]" pageId="16" pageNumber="17">
<taxonomicName id="EB8970EAF059FFE5FF1EFBED7EDAFB84" ID-CoL="4FVZ3" authority="Ebert, 1965" authorityName="Ebert" authorityYear="1965" box="[151,428,1103,1127]" class="Insecta" family="Geometridae" genus="Phaselia" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="erika">
<emphasis id="1EFDD77BF059FFE5FF1EFBED7E5DFB84" box="[151,299,1103,1127]" italics="true" pageId="16" pageNumber="17">Phaselia erika</emphasis>
<materialsCitation id="9CE10134F059FFE5FEBBFBED7EDAFB84" box="[306,428,1103,1127]" collectionCode="SMNK" collectorName="Stuttgarter Beitrage &amp; Naturkunde. Nr." country="Afghanistan" location="Sarobi" pageId="16" pageNumber="17" specimenCount="1" typeStatus="holotype">
<bibRefCitation id="48187698F059FFE5FEBBFBED7EDAFB84" author="Ebert, G." box="[306,428,1103,1127]" pageId="16" pageNumber="17" pagination="1 - 32" refId="ref30159" refString="Ebert, G. (1965) Afghanische Geometriden (Lep.) I. Stuttgarter Beitrage zur Naturkunde, 142, 1 - 32." type="journal article" year="1965">Ebert, 1965</bibRefCitation>
</materialsCitation>
</taxonomicName>
. 
<collectorName id="817C6EBFF059FFE5FE3FFBED7D08FB84" box="[438,638,1103,1127]" pageId="16" pageNumber="17">Stuttgarter Beiträge</collectorName>
zur 
<collectorName id="817C6EBFF059FFE5FD25FBED7C24FB84" box="[684,850,1103,1127]" pageId="16" pageNumber="17">Naturkunde. Nr.</collectorName>
142: 17 pl. 1, fig. 15; pl. 3 figs 8, 9. 
<typeStatus id="F332B5CBF059FFE5FB47FBED7A5AFB84" box="[1230,1324,1103,1127]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
♁, SMNK (
<collectingCountry id="549E4BF9F059FFE5FF47FBCC7E38FB65" box="[206,334,1134,1158]" name="Afghanistan" pageId="16" pageNumber="17">Afghanistan</collectingCountry>
, 
<location id="29565DB2F059FFE5FED1FBCC7EE9FB65" LSID="urn:lsid:plazi:treatment:A420BA7FF059FFE5FF1EFC447F9BFAD6:29565DB2F059FFE5FED1FBCC7EE9FB65" box="[344,415,1134,1158]" country="Afghanistan" name="Sarobi" pageId="16" pageNumber="17">Sarobi</location>
).
</paragraph>
<paragraph id="2C360B69F059FFE5FF1EFB0C7F9BFAD6" blockId="16.[151,1436,1198,1333]" pageId="16" pageNumber="17">
<emphasis id="1EFDD77BF059FFE5FF1EFB0C7EF5FB2A" bold="true" box="[151,387,1198,1225]" pageId="16" pageNumber="17">Taxonomic remark.</emphasis>
The results of the examination of the 
<typeStatus id="F332B5CBF059FFE5FCBAFB0D7C15FB2A" box="[819,867,1199,1225]" pageId="16" pageNumber="17">type</typeStatus>
materials and a large number of specimens in this study suggest that the species 
<taxonomicName id="EB8970EAF059FFE5FE6DFB717D4FFB0E" ID-CoL="4FVZ3" authorityName="Ebert" authorityYear="1965" box="[484,569,1235,1261]" class="Insecta" family="Geometridae" genus="Phaselia" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="erika">
<emphasis id="1EFDD77BF059FFE5FE6DFB717D4FFB0E" box="[484,569,1235,1261]" italics="true" pageId="16" pageNumber="17">P. erika</emphasis>
</taxonomicName>
can be subdivided into at least three allopatric subspecies which are separated from each other by large geographic distances. The regions between the ranges of these subspecies need further studies.
</paragraph>
</subSubSection>
</treatment>
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