Five new Palaearctic species of Docosia (Diptera: Mycetophilidae), with updated molecular phylogeny of the genus Author Ševčík, Jan 639373EF-6D46-40F6-929B-0C861C6C39F6 University of Ostrava, Faculty of Science, Department of Biology and Ecology, Chittussiho 10, CZ- 710 00 Ostrava, Czech Republic. Silesian Museum, Nádražní okruh 31, CZ- 746 01 Opava, Czech Republic. Institute of Agricultural and Environmental Sciences, Estonian University of Life Sciences, Kreutzwaldi st 5 D, 51006 Tartu, Estonia. sevcikjan@hotmail.com Author Burdíková, Nikola F7CCFBF7-CB4E-489A-95AC-7346268EB0E0 University of Ostrava, Faculty of Science, Department of Biology and Ecology, Chittussiho 10, CZ- 710 00 Ostrava, Czech Republic. Silesian Museum, Nádražní okruh 31, CZ- 746 01 Opava, Czech Republic. Institute of Agricultural and Environmental Sciences, Estonian University of Life Sciences, Kreutzwaldi st 5 D, 51006 Tartu, Estonia. burdikova@seznam.cz Author Kaspřák, David 665C08DD-1099-4F88-B528-A9441AFC6A36 University of Ostrava, Faculty of Science, Department of Biology and Ecology, Chittussiho 10, CZ- 710 00 Ostrava, Czech Republic. Silesian Museum, Nádražní okruh 31, CZ- 746 01 Opava, Czech Republic. Institute of Agricultural and Environmental Sciences, Estonian University of Life Sciences, Kreutzwaldi st 5 D, 51006 Tartu, Estonia. davidkasprak@gmail.com Author Kurina, Olavi FB595938-73A2-4DBC-9ABB-77E81D13DFE1 Institute of Agricultural and Environmental Sciences, Estonian University of Life Sciences, Kreutzwaldi st 5 D, 51006 Tartu, Estonia. olavi.kurina@emu.ee text European Journal of Taxonomy 2020 2020-09-17 717 3 26 journal article 20950 10.5852/ejt.2020.717.1095 b9fb54f7-1883-4a6d-99b4-76f855a4d1cd 4043697 1358E5B2-056C-4DF2-9C4E-0189E1086712 Docosia peloponnensis Ševčík sp. nov. urn:lsid:zoobank.org:act: 979DF75A-E677-4C63-8B13-BB3F3270115F Figs 4 , 7D Differential diagnosis This species is unique among the species of Docosia in having a simple gonostylus with a sickle-like posterior projection. Etymology The species name refers to the type locality. Type material Holotype GREECE • ♂ ; Peloponnese , Valtesiniko env.; 1100 m a.s.l. ; 22 Apr. 2016 ; J. Ševčík leg.; sweeping along small brook; in a pinned microvial with glycerol; specimen after DNA extraction; GenBank: MT293011 , MT293042 , MT293059 , MT292993 ; NMPC- JSDO15. Paratype GREECE • 1 ♂ ; Peloponnese , Valtesiniko env.; 1100 m a.s.l. ; 26 Apr. 2016 ; M. Tkoč leg.; sweeping along small brook ; in a pinned microvial with glycerol; JSL-UOC-JSDO15b. Description Male (n = 2) MEASUREMENTS. Length of wing 4.2 mm . HEAD. Blackish brown with numerous pale setae. Three ocelli, lateral ones almost touching compound eyes, separated from eye margins by less than half of their own diameter. Clypeus blackish, with setae pale. Mouthparts light brownish. Palpus brownish yellow, basal segment and apical part of last palpomere darker. Scape, pedicel and all flagellomeres dark brown. Flagellomeres cylindrical, flagellomeres 1 to 7 about twice as long as broad, distal flagellomeres (8 to 14) slightly conical, 2.5 to 3 times as long as broad. THORAX.All parts of thorax blackish brown, with light setae. Scutellum with numerous setae and several submarginal pale bristles about twice as long as scutellum. Antepronotum and proepisternum with pale bristles and short darker setae. Laterotergite and other pleural parts bare. Haltere pale yellow. LEGS. All coxae yellow, with basal third darkened. Femora mostly yellow, fore and mid femur dark along ventral margin, more extensively in proximal third; similarly hind femur, brownish also around tip. All trochanters blackish brown. Tibiae and tarsi yellow, tarsal segments seemingly brownish because of dense setulae. Fore tibia apicomedially with semicircular tibial organ (anteroapical depressed area), without strong setae, only densely covered with fine setulae. Fig. 4. Docosia peloponnensis Ševčík sp. nov. , ♂, terminalia. A . Lateral view. B . Posterior view. C . Ventral view of gonostylus. D . Dorsal view of cerci. E . Dorsal view of tergite 9. F. Ventral view of aedeagal complex. G . Lateral view of aedeagal complex. Scale bars = 0.1 mm. WINGS. Hyaline, unmarked. Radial veins and r-m dark brown, other veins paler, basal parts of m faint, almost not traceable. Sc, Rs and basal third of cu-stem asetose, other veins setose. Costa reaches to 0.45 of distance between R 5 and M 1 . Sc ends in R slightly beyond level of beginning of m-stem. Posterior fork begins before anterior fork, approximately at level of basal third of r-m. ABDOMEN. All dark brown. TERMINALIA ( Figs 4 A–E, 7D). Dark brown except lighter gonostyli and posterior margin of gonocoxites. Tergite 9 about as long as broad, slightly narrowing towards base, with slightly concave posterior margin. Posteroventral margin of gonocoxites without lateral projections, with distinct U-shaped medial incision and narrow digitiform medioventral process. Gonostylus simple, slightly bulging medially, with aggregation of small setae at mediobasal corner and with characteristic sickle-like posterior projection, with prominent basal setae at its ventral and dorsal sides. Cercus with 14 combs of retinacula. Female Unknown. Biology Unknown.