Paleocene origin of the cockroach families Blaberidae and Corydiidae: Evidence from Amur River region of Russia Author Vršanský, Peter Author Vidlička, Ľubomír Author Barna, Peter Author Bugdaeva, Eugenia Author Markevich, Valentina text Zootaxa 2013 3635 2 117 126 journal article 10.11646/zootaxa.3635.2.2 d4675d41-094b-402d-a545-04c1a927fe3b 1175-5326 219088 B74B0B78-911E-41FE-879F-7D41C8A77DBF Corydiidae Saussure, 1864 (= Polyphagidae Walker, 1868 ) Corydiinae Saussure, 1864 (= Polyphaginae Walker, 1868 ) Corydiini sensu Rehn, 1951 Diagnosis (after Rehn 1951). Both sexes with at least tegmina present, wings usually present, but sometimes considerably reduced. Tegmina varying from normal to somewhat reduced, obovate and densely coriaceous (mostly in females). Humeral area more developed than in Polyphagini, if coriaceous then broadly expanded. Sc rami regular, not crowded. R without posterior branches, most branches terminating anteriorly, some apically, instead of curving posteriorly. M with free base, its branching regular and direct. Cu not curving distinctly away from plical furrow, CuP not joining cubitus. Diagnosis (after Walker 1868). Female: Body short-elliptical, convex, dull, very thickly and minutely punctured. Head shining, impressed between the eyes, with a transverse furrow near the mouth. Eyes not far apart. Second joint of the palpi subclavate; third slightly securiform, very much longer than the second. Antennae setaceous, submoniliform, not more than half the length of the body; first, second and third joints short; following joints very short. Prothorax extending somewhat beyond the head and over the basal part of the fore wings when they are expanded, rounded in front and on each side, slightly furrowed along each side; its breadth along the hind border more than twice its length; hind border hardly rounded; hind angles slightly falcate; a lyre-shaped mark in the disk. Mesothorax, metathorax, pectus and abdomen shining, mostly smooth. Abdomen with the segments above and beneath near the tip retracted in the middle towards the disk; sides fringed, with bristles; subanal lamina small, bilobed. Cerci lanceolate, submoniliform, setose. Legs stout; tibiae armed with some strong spines; first joint of the tarsi twice the length of the fifth, which is very much longer than the second. Fore wings coriaceous, membranous towards the border; costa much rounded; tips conical; principal veins distinct in the coriaceous part; transverse sectors numerous, irregular. Hind wings membranous, strongly and thickly reticulated; transverse sectors numerous, irregular. Type species . Ergaula carunculigera (Gerstaecker, 1861) Composition (updated from Princis 1963). Ergaula Walker, 1868 = Dyscologamia Saussure, 1893 ( type is cesticulata = pilosa ) = Parapolyphaga Chopard, 1929 ( type is erectipilis = pilosa )?= Netherea Vršanský et Anisyutkin, 2004 ( type is haatica ) Ergaula capensis (Saussure, 1893) ( Nigeria , Cameroon , Democratic Republic of the Congo , Congo , Uganda , Kenya , Tanzania , Zambia , Zimbabwe , Angola ) = Dyscologamia capensis Saussure, 1893 = Dyscologamia wollastoni Kirby, 1909 Ergaula capucina (Brunner von Wattenwyl, 1893) ( Myanmar ) = Homoeogamia capucina Brunner von Wattenwyl, 1893 Ergaula carunculigera (Gerstaecker, 1861) ( Philippines (Luzon)) = Corydia carunculigera Gerstaecker, 1861 = Ergaula scaraboides Walker, 1868 Ergaula funebris (Hanitsch, 1933) (Borneo) = Dyscologamia funebris Hanitsch, 1933 Ergaula nepalensis (Saussure, 1893) ( Nepal , Myanmar ) = Dyscologamia nepalensis Saussure, 1893 Ergaula pilosa (Walker, 1868) (Sumatra, Malaysia , Java, Borneo) = Zetobora pilosa Walker, 1868 = Dyscologamia cesticulata Saussure, 1893 = Dyscologamia chopardi Hanitsch, 1923 = Parapolyphaga erectipilis Chopard, 1929 = Polyphaga sumatrensis Shelford, 1908 Ergaula silphoides (Walker, 1868) ( Cambodia ) = Polyphaga silphoides Walker, 1868 Ergaula atica Vršanský et Anisyutkin, 2008 ( Israel ) extinct,?Eocene (based on male)?= Netherea haatica Vršanský et Anisyutkin, 2008 ( Israel ) extinct,?Eocene (based on female) Ergaula spp. ( Germany ) extinct, Eocene (common in Messel, based on both sexes)