Phylogenetic and morphological resolution of the Helobdella stagnalis species-complex (Annelida: Clitellata: Hirudinea) Author Saglam, Naim Author Kutschera, Ulrich Author Saunders, Ralph Author Saidel, William M. Author Balombini, Katherine L. W. Author Shain, Daniel H. text Zootaxa 2018 2018-04-03 4403 1 61 86 journal article 30344 10.11646/zootaxa.4403.1.3 82592d82-9f4b-4f22-8c93-ca00c436d13b 1175-5326 1212238 7B575B1A-9158-46CD-B275-5A3224693663 Helobdella eriensis n. sp. ( FPWS 2-Farm Pond) ( Figs. 2 D-7D) ( Holotype : ANSP GI 19494) (LSID: urn:lsid:zoobank.org:act:91CD93B2-3B58-42C9-B640-9C92FD8827CC )— Dorsal body light to yellowish-grey, sometimes with greenish tint. Ventral same as H. serendipitious n. sp. Total of 67 annul; I, II, III uniannulate, IV, V biannulate, VI-XXIII triannulate, XXV biannulate, XXVI-XXVII uniannulate. One pair of small, elliptical eyes clearly separated on segment III ( Fig. 7D ). Male and female gonopores between XI a2/a3 (24th/25th annuli) and XIa3/XIIa1 (25th/26th annuli), respectively; clitellum between XI-XIIth segments. Six pairs of testisacs intersegmentally arranged, located as in H. modesta . Ejaculatory bulbs similar to H. stagnalis , Ovisacs in XII segment large, connected with long and curled common oviduct. Oviduct shorter than ovisac ( Figs. 2D , 4D ). Six pairs of crop caeca, 6th directed backwards and short extending up to fourth intestinal caecum. Four pairs of intestinal caecum, first three curved slightly upward and fourth curved downward. Rectum is long with twisted “S” shape ( Figs. 3D , 5D ).