Order Rodentia - Family Muridae Author Wilson, Don E. Author Reeder, DeeAnn text 2005 The Johns Hopkins University Press Baltimore Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 2 1189 1531 book chapter 0-8018-8221-4 10.5281/zenodo.7316535 Dipodillus (Petteromys) campestris (Loche 1867) [Gerbillus] campestris Loche 1867 , Expl. Sci. Alg. Zool. Mamm.: 106 . Type Locality: Algeria , Constantine Province , Philipeville. Vernacular Names: North African Dipodil . Synonyms: Dipodillus (Petteromys) brunnescens ( Ranck 1968 ) ; Dipodillus (Petteromys) cinnamomeus Cabrera 1922 ; Dipodillus (Petteromys) dodsoni Thomas 1913 ; Dipodillus (Petteromys) gerbii (Loche 1867) ; Dipodillus (Petteromys) haymani (Setzer 1958) ; Dipodillus (Petteromys) hilda (Thomas 1918) ; Dipodillus (Petteromys) minutus (Loche 1867) ; Dipodillus (Petteromys) patrizii de Beaux 1932 ; Dipodillus (Petteromys) quadrimaculatus (Lataste 1882) ; Dipodillus (Petteromys) riparius Cabrera 1922 ; Dipodillus (Petteromys) rozsikae Thomas 1908 ; Dipodillus (Petteromys) somalicus Thomas 1910 ; Dipodillus (Petteromys) venustus (Sundevall 1843) ; Dipodillus (Petteromys) wassifi Setzer 1958 . Distribution: N Africa, from Morocco to Egypt and Sudan . Conservation: IUCN – Critically Endangered as Gerbillus quadrimaculatus , Lower Risk (lc) as G. campestris . Discussion: Subgenus Petteromys . Nineteen species-group names have been associated with D. campestris by different authors in various combinations, as summarized by Lay (1983) , who also noted that most opinions lacked supportive evidence and that some of the synonyms are unidentifiable or nomina nuda . Lay (1983) remarked that although most authors followed Ellerman and Morrison-Scott (1951) in listing quadrimaculatus equivocally under G. nanus , it should be kept separate until a revision is available. Pavlinov et al. (1990) synonymized it with Dipodillus campestris and we follow their allocation. Musser and Carleton (1993) had allocated hilda to G. nanus , but it belongs here (Aulagnier, in litt.). Those names listed here are probably correctly associated with D. campestris , but the species requires refined definition through careful systematic revision. Different geographical populations were reviewed by Aulagnier and Thevenot (1986 , Morocco ) Kowalski and Rzebik-Kowalska (1991 , Algeria ), Ranck (1968 , Libya ), and Osborn and Helmy (1980 , Egypt ). Benazzou and Zyadi (1990) conducted a biometric study analyzing variation among Moroccan populations. Dobigny et al. (2001 a, b ) documented chromosomal traits (2n = 56, FNa = 68) from Adrar des Iforas in NE Mali , which are similar to karyotypes reported from the Maghreb and S Niger ( Dobigny et al., 2002 b ), and from Tunisia ( Chetoui et al., 2002 ). Significance of APRT gene sequences in illuminating substitution rate variation among D. campestris and other muroids presented by Fieldhouse et al. (1997) . Cockrum and Setzer (1976:643) clarified the author and date of publication of campestris (also see Kowalski and Rzebik-Kowalska, 1991 ). Reviewed by Pavlinov et al. (1990) . Fragments identified as D. campestris are reported from probable middle Pleistocene sites in Morocco ( Amani and Geraads, 1993 ; Geraads, 1994 ), and D. bibersoni was described (as Gerbillus ) from late Pliocene fossils in Moroccan strata and is considered closely related to D. campestris ( Geraads, 1995 ) .